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Evolutionary Systematics. 5 2021, 193-261 | DOI 10.3897/evolsyst.5.63435 


> PENSUFT. 


EVOLUTIONARY 
SYSTEMATICS 


An illustrated catalogue of the type specimens of Lepidoptera 
housed in the Zoological Museum Hamburg (ZMH): 


Part I. superfamily Papilionoidea 


Reza Zahiri!:*, Vazrick Nazari?, Hossein Rajaei*, Martin Wiemers?, 
Maryam Fatahi', Matthias Seidel’, Thure Dalsgaard', Martin Husemann! 


oF WN BP 


http://zoobank. org/984E 15D8&-80E0-4B7D-A84F-92BB0AD4EA 73 


Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany 

Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., KLA 0C6, Ottawa, Ontario, Canada 
Independent Researcher, P.O. Box 45063, Ottawa, Ontario K2M 2Y1, Canada 

Department Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany 

Department of Systematics and Biogeography, Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Miincheberg, Germany 


Corresponding author: Reza Zahiri (reza.zahiri@inspection.gc.ca, reza.zahiri@gmail.com) 


Academic editor: Danilo Harms # Received 22 January 2021 # Accepted 10 June 2021 # Published 20 August 2021 


Abstract 


We provide an updated catalogue of the type material of the lepidopteran superfamily Papilionoidea deposited in the Zoological Mu- 
seum of Hamburg (ZMH). We report 414 specimens labelled as “types” belonging to nine species (all of valid names), 74 subspecies 
(44 valid names and 30 synonyms), 59 invalid infrasubspecific names under the ICZN code, and 23 specimens of 16 “in litteris” 
(= unavailable) names. Out of the 414 specimens labelled as “types”, 171 specimens are primary types (8 holotypes/lectotypes and 
163 syntypes) and 80 are secondary types; 120 specimens are infrasubspecific and hence invalid and are considered as “original 
specimens”; and 43 specimens are treated here as “Non-type” specimens (topotype). We present a full bibliography of the original 


descriptions and illustrations for all of these taxa, aiming to provide a comprehensive taxonomic guide to this collection. 


Key Words 


Biodiversity, CeNak, Herbert Weidner, ZMH, digitization, Georg Warnecke, Hanan Bytinski-Salz 


Introduction 


Natural history collections are the main archives of 
biodiversity. They focus on collecting, maintaining and 
documenting natural specimens, a crucial task, especially 
in times of global biodiversity decline (Dirzo et al. 2014). 
The Zoological Museum of Hamburg (ZMH) holds large 
collections across all animal groups. With approximately 
five million specimens and several thousand primary 
types, the entomological collection is among the most 
important natural history archives in Germany. Yet, the 
collections are overall not well documented and updated 
type catalogues are largely lacking. 

The last ZMH Lepidoptera catalogue was published 
by Herbert Weidner, a former curator of the collection, 


in 1974 (Weidner 1974). Since the publication of this 
work more than 40 years ago, no further types have been 
documented. We initiated a comprehensive effort to doc- 
ument all of the type material preserved in the ZMH in 
English language (Sartori et al. 2016; Dey and Husemann 
2018a, b; Harms and Dupérré 2018; Monod et al. 2019; 
Henningsen et al. 2020; Simoes et al. 2021; Zahiri et al. 
2021). We performed a detailed inventory throughout the 
collection. As a part of this inventory, the entire collec- 
tion was thoroughly searched and types were transferred 
to a separate, specially protected type collection. All type 
specimens were compared with the original descriptions, 
photographed, databased and relabeled, if required. The 
original high-quality photographs are available online 
through the FUNDus! Portal of the University of Ham- 


Copyright Reza Zahiri et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits 
unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 


194 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


burg (https://www.fundus.uni-hamburg.de/). The com- 
plete data of the type material is provided in two table 
formats: taxa format (Suppl. material 1: Table S1) and 
specimen format (Suppl. material 2: Table S2). Our first 
Lepidoptera catalogue (Zahiri et al. 2021) on the super- 
families Hepialoidea, Cossoidea and Zygaenoidea has 
resulted in an updated list of 259 as “type” labelled spec- 
imens belonging to 13 valid species, 34 subspecies (16 
valid and 18 invalid names), and 36 names of invalid in- 
frasubspecific rank (based on the ICZN code). 


Material and methods 
Photography 


All images were taken with a custom-made digital im- 
aging system (DUN Inc.) with stacking capabilities. The 
images where either taken with a Canon EOS S5DSR with 
a 65 mm or 100 mm lens and stacked with Zerene Stacker 
(PMax algorithm) or with a Canon EOS 6D with a 55 mm 
or 65 mm lens and stacked with Helicon Focus 5.3. 


Taxonomy 


All taxonomic changes were made after studying the rele- 
vant original descriptions of the type material. Synonyms 
that have already been proposed are listed only if discussed 
and the type material was examined. Classification used 
herein follows van Nieukerken et al. (2011). The high- 
er taxonomy of the families Papilionidae, Hesperiidae, 
Pieridae, Lycaenidae, Riodinidae, and Nymphalidae, as 
well as the classification below the family level presented 
here, are based on a combination of recent comprehen- 
sive morphological and molecular studies (Paulus et al. 
1996; Wahlberg et al. 2009; Heikkila et al. 2011; Kawa- 
hara et al. 2014), as well as the updated checklist of the 
European butterflies (Wiemers et al. 2018). Finally, like 
almost all major Lepidoptera databases and catalogues, 
we did not follow the principle of gender agreement in 
species names (Sommerer 2002). The recommendation 
not to follow the articles 31.2 and 34.2 of the ICZN code 
was adopted at the General Meeting of the Societas Euro- 
paea Lepidopterologica (SEL) in a resolution proposed at 
the 13" European Congress of Lepidopterology in Korsor 
(Denmark) on June 4, 2002 (van Nieukerken 2019). 


Type material 


Type localities are cited in their original spelling or with 
additional remarks (in square brackets), if required. All 
data was transcribed verbatim as appearing on the respec- 
tive labels. For unreadable data a question mark (?) was 
used. Double slash (//) separates data on different labels, 
a single slash (/) separates lines within each label, and 
semi-colon (;) separates different specimens. Additional 
comments are placed in square brackets. 


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The ICZN (1999) terminologies (e.g., holotype, syn- 
type, lectotype, neotype, paratype, paralectotype, al- 
lotype, cotype etc.) are available at the ICZN website 
(http://iczn.org/content/Glossary) and ICZN articles cited 
in the text can be found below: 


Infrasubspecific taxon — a taxon at lower rank than that 
of subspecies. The names of such taxa are not regu- 
lated by the Code. 

Article 45.6.1. — A name is infrasubspecific if its author 
expressly gave it infrasubspecific rank, or if the con- 
tent of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity; 

Article 45.6.2. — A name 1s deemed to be infrasubspe- 
cific if its author used one of the terms “aberration”, 
“ab.” or “morph”. 

Article 45.6.3. — A name is deemed to be infrasubspecific 
if it was first published after 1960 and the author ex- 
pressly used one of the terms “variety” or “form” (in- 
cluding use of the terms “var.”, “forma”, “v.” and “f.”’): 

Article 45.6.4. — A name is subspecific if first published 
before 1961 and its author expressly used one of the 
terms “variety” or “form” (including use of the terms 
“var.”, “forma”, “v.” and “f.”’), unless its author also 

expressly gave it infrasubspecific rank, or the con- 

tent of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity, in which 
case it is infrasubspecific. 


Type specimens deposited at the 
Zoological Museum Hamburg (ZMH) 


Superfamily Papilionoidea Latreille, 1802 


Papilionoidea (butterflies), currently containing seven 
families with approximately 18,800 described species 
worldwide (van Nieukerken et al. 2011), is one of the 
well-studied lepidopteran groups. The family Nymphali- 
dae (Brush-footed butterflies) with 6,152 described species 
is the most diverse taxonomic group followed by Lycaeni- 
dae (Blues and coppers) with 5,201 species, Hesperiidae 
(Skippers) with 4,113 species, Riodinidae (Metalmark 
butterflies) with 1,532 species, Pieridae (Whites or Yel- 
low-whites) with 1,164 species, Papilionidae (Swallotails) 
with 570 species and Hedylidae with only 36 species (van 
Nieukerken et al. 2011). 


Family Papilionidae Latreille, 1802 
Subfamily Parnassiinae Duponchel, 1835 


1. tridentina Dannehl, 1925 


Original combination. “Parnassius apollo L. (tridentina 
Dannehl)” Dannehl, 1925 Ent. Z. 39: 6. 

Current combination. Parnassius apollo tridentina 
Dannehl, 1925. 

Current status. Valid subspecies. 


Evolutionary Systematics 5 2021, 193-261 


Type material. Syntypes 24'¢ (ZMH 61579-61580) 
(Fig. 1). “Teriol. Mer. / Sum. alp. / Palon / 1.7.24 Ibor 
[?] / Dannehl” // “Cotype” // “1.7. 1924 / Mt. Palon 
/ Stidtirol / Dannehl” // “f. tridentina Dhl.” // “Mus. 
Altona comm / Eing Nr 3-65” // “ZMH 61579”; “10.6” 
// “Dannehl / Teriol. mer. / Alp. vai. / Mt. Palon / 1400 
m.” // “Cotype” // “10.6 / Mt. Palon / Stidtirol / Dan- 
nehl” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 
61580”. 

Type locality. Italy: “Lokalrasse aus den ostlichen 
Teilen der Trientiner (Sarcataler) Alpen; westl. Etschufer 
sudlich Trient. 1200—1600 m.” 


2. mendolensis Dannehl, 1925 


Original combination. “Parnassius apollo L. (mendo- 
lensis Dannehl).” Dannehl, 1925 Ent. Z. 39: 6. 

Current combination. Parnassius apollo mendolen- 
sis Dannehl, 1925. 

Current status. Valid subspecies. 

Type material. Syntypes 1419 (ZMH 61581-61582) 
(Fig. 2). “mendolensis 2” // “Teriol mer. / Mendel / 24. 
juni / coll. F. Dannehl” // “Cotype” // “Mus. Altona 
comm / Eing Nr 3-65” // “ZMH 61581”; “mendolensis 
3” // “Teriol mer. / Mendel / 3.Juni.22 / coll. F. Dannehl” 
// “Cotype” // “Cotype” // “Mus. Altona comm / Eing Nr 
3-65” // “ZMH 61582”. 

Type locality. Italy: “Flegt auf den oberen der Men- 
delstrafe bei Bozen und in den meisten ostlichen Talein- 
schnitten des Mt. Roen, 500-1800 m.” 


3. sevensis Kesenheimer, 1921 


Original combination. “Parn. apollo Linné, nov. subsp. 
sevensis Ksh.” Kesenheimer, 1921 Ent. Z. 34: 63. 

Current combination. Parnassius apollo sevensis 
Kesenheimer, 1921. 

Current status. Junior subjective synonym of Par- 
nassius apollo meridionalis Pagenstecher, 1909. 

Type material. Syntypes 1412 (ZMH 61583- 
615584) (Fig. 3). “Cotype” // “apollo / sevensis / Cotype 
9 / 19.7.14 / Aalfeld — Sternsee / H.W. Kesenheimer” 
// “Cotype” // “Sammlung / G. Warnecke / Eing Nr. 5, 
1949” // “ZMH 61583”; “Sammlung / G. Warnecke / 
Eing Nr 51949” // “apollo sevensis / Cotype @ / 19.3.14 
/ Aalfeld - Sternsee / H.W. Kesenheimer” // “Cotype” // 
“Cotype” // “ZMH 61584”. 

Type locality. France: “von Sewen im Masmiunstertal 
in den Vogesen”. 


4. kricheldorffi Eisner, 1928 


Original combination. “Parnassius apollo, forma 
kricheldorffi (m.)” Eisner, 1928 Int. Ent. Z. 22: 309. 

Current combination. Parnassius apollo krichel- 
dorffi Eisner, 1928. 


195 


Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 61585) (Fig. 4). 
“Asturien / Picos de Europa / 1800-2000 m. / 20. —28. 
VII.1928 / leg. A. Kricheldorff” // “v. Kricheldorfi Eisn.” 
// “Co-Type / e Collection / Bang Haas” // “Cotype” // 
“Parnassius V. / Kricheldorffi Es. / det. Eisner” // “Sam- 
mlung / G Warnecke / Eing Nr.5. 1949” // “ZMH 61585”. 

Type locality. Spain: “Asturien, Picos de Europa, 
1800—2000 m”. 

Remarks. Eisner (1928) proposed this name as a form 
of P. apollo (Linnaeus, 1758). According to article 45.6.4 
(ICZN 1999), it is subspecific if first published before 
1961 and the author expressly used the terms “variety” 
or “form”. 


5. ab. praetexta Gelpke, 1929 


Original combination. “Parnassius apollo L. f. valesi- 
acus ab. praetexta m.” Gelpke, 1929 Int. Ent. Z. 23: 123. 

Current combination. Parnassius apollo valesiacus 
ab. praetexta Gelpke, 1929. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 12 (ZMH 
61586) (Fig. 5). “Type / P. apollo L: valesiacus / Fruhst. 
/ ab. praetexta Gelpke / I. E. Z. Guben 23 Jahrg 1929 / 
S. 122 [illegible]” // “13.7. 1927 / Lotschental / ob Wiler 
/ Wallis 1550 m” // “Type” // “ab. praetexta Gelpke” // 
“Mus. Altona comm / Eing. Nr. 3-65 // ZMH 61586”. 

Original locality. Switzerland: Swiss Alps. “oberhalb 
Wiler”. 

Remarks. Gelpke (1929) proposed this name as an ab- 
erration of P. a. valesiacus Fruhstorfer, 1906. Therefore, 
as stated by articles 45.6.1 and 45.6.2 ICZN 1999) it is 
deemed to be an infrasubspecific name (if the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 


6. ab. uniformis Stauder, 1922 


Original combination. “Parnassius apollo ab. uniform- 
is” Stauder, 1922 Soc. Ent. 36: 10 (Note) 

Current combination. Parnassius apollo ab. uni- 
formis Stauder, 1922. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
61587) (Fig. 6). “Unter-Italien / Stauder 1920” // “uni- 
formis Stdr / Type” // “ab. uniformis Stauder / Type” // 
“Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 
61587”. 

Original locality. Italy: “Unteritalien”. 

Remarks. Stauder (1922) proposed this name as an 
aberration of P. apollo (Linnaeus, 1758). Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author used one of the terms 
“aberration”, “ab.” or “morph”) and is hence unavailable. 


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196 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


' Mus. Aitona comm 
Eing Ne 3-65 


| Mus. Aitona comm’ 
Eing ‘Nre 3- 65° 


z 


ZMH - 61582 


i ae i 
er | 
. 


Mus. Aitena comm 
2 
Fing, Neo 3- 0d 


Plate 1. 1. Parnassius apollo tridentina Dannehl, 1925; 2. Parnassius apollo mendolensis Dannehl, 1925; 3. Parnassius apollo 
sevensis Kesenheimer, 1921; 4. Parnassius apollo kricheldorffi Eisner, 1928; 5. Parnassius apollo valesiacus ab. praetexta Gelpke, 
1929: 6. Parnassius apollo ab. uniformis Stauder, 1922. a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


7. f. bispupillata Turati, 1918 


Original combination. “Parnassius apollo pumilus 
Stich. forma bispupillata” Turati, 1918 Atti. Soc. Ital. Sci. 
Nat. 57: 40. 

Current combination. Parnassius apollo pumilus f. 
bispupillata Turati, 1918. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 14 (ZMH 
61588) (Fig. 7). “Calabrien / [illegible], 1800 m / Stauder, 
5.7.20” // “bispupillata” // “bispupillata” // “Cotype” // 
“Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 
61588”. 

Original locality. Italy: Calabria. 

Remarks. Turati (1918) proposed this name as a form 
of P. a. pumilus Stichel, 1906. Therefore, as stated by arti- 
cle 45.6.1 (ICZN 1999) it is deemed to be an infrasubspe- 
cific name (if the author expressly gave it infrasubspecific 
rank) and is hence unavailable. 


8. f. appendiculata Turati, 1918 


Original combination. “Parnassius apollo pumilus 
Stich. forma appendiculata (f. nuova)” Turati, 1918 Atti. 
Soc. Ital. Sci. Nat. 57: 41. 

Current combination. Parnassius apollo pumilus f. 
appendiculata Turati, 1918. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 1411 9 (ZMH 
61589-61590) (Fig. 8). “Calabrien / Aspermonte, 1800 m 
/ Stauder 5.7.1920” // “flavomaculata / appendiculata” // 
“flavomaculata / appendiculata” // “Sammlung / G. War- 
necke / Eing. Nr.5, 1949” // “Cotype” // “ZMH 61589”; 
“Calabrien / Aspromonte, 1800 m / Stauder 5.7.1920” // 
“nigricans / appendiculata” // “9 nigricans / appendicu- 
lata” // “Sammlung / G. Warnecke / Eing. Nr.5, 1949” // 
“Cotype” // “ZMH 61590”. 

Original locality. Italy: Calabria. 

Remarks. Turati (1918) proposed this name as a form 
of P. a. pumilus Stichel, 1906. Therefore, as stated by arti- 
cle 45.6.1 (ICZN 1999) it is deemed to be an infrasubspe- 
cific name (the author expressly gave it infrasubspecific 
rank) and is hence unavailable. 


9. ab. lacrimans Marschner, 1911 


Original combination. “Parnassius apollo L. ab. lacri- 
mans” Marschner, 1911 Dt. Ent. Z. Iris 25: 131. 

Current combination. Parnassius apollo ab. lacri- 
mans Marschner, 1911. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
61591) (Fig. 9). “Schweizer Jura / Bozinger Berg / Juli 


197 


1919/ecoll Marschner / coll. Dr. Gelpke / 1942” // “Type 
/ Parn. apollo L. / f. lacrimans / Marschner / Collect. H. 
Marschner” // “ab. lacrimans Marschn.” // “Typen” // 
“Mus Altona comm. / Eing Nr. 3 05” // “ZMH 61591”. 
Original locality. Switzerland: “schweizer Jura”. 
Remarks. Marschner (1911) proposed this name as 
an aberration of P. apollo (Linnaeus, 1758). Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author used one of the terms 
“aberration”, “ab.” or “morph”) and is hence unavailable. 


10. oenipontanus Stauder & Felkel, 1921 


Original combination. “Parnassius apollo oenipontanus, 
n. subsp.” Stauder & Felkel, 1921 Ent. Anz. 1: 123-124. 

Current combination. Parnassius apollo oeniponta- 
nus Stauder & Felkel, 1921. 

Current status. Valid subspecies. 

Type material. Syntypes 104.4 (ZMH 61592-61601) 
(Fig. 10). “el. / 9./6.1924 / [illegible] / Innsbruck” // 
“Coll. Janiter / Leihgabe der / Jungiusgesellschaft / Eing 
Nr. 13, 1958” // “ZMH 61592”; “Type” // “Innsbruck / 
10.6.1913 / J. Felkel” // “oenipontanus Stder / Cotype” // 
“Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 
61593”; “Type” // “Innsbruck Stilltal” // “Silltal /7.6.1921 
/ J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlu- 
ng / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61594”; 
“Cotype” // “oenipontanus / Stauder — Felkel / Innsbruk” 
// “Silltal / 23.5.1921 / J. Felkel” // “Coll. G. Warnecke / 
Eing. Nr. 11,1983” // “ZMH 61595”; “Type” // “Silltal / 
19.5.1921 / J. Felkel” / “oenipontanus / Stauder — Felkel” 
/“Tnnsbruck / Silltal” // “oenipontanus Stder / Cotype” // 
“Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 
61596”; “Cotype” // “Innsbruck / gebirge / 21/6/1914 / J. 
Felkel leg.” // “oenipontanus Stder / Cotype” // “Sammlu- 
ng / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61597”; 
“Type” // Innsbruck / Silltal” // “Silltal / 19.5.1921 / J. 
Felkel” // “oenipontanus Stauder / Cotype” //“Summtng / 
G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61598”; “Type” 
// “H. Stauder / Nordtirol / Patsch / 11/6/1920” // “oeni- 
pontanus Stauder / Cotype” // “Sammlung / G. Warnecke 
/Eing. Nr. 5,1949” // “ZMH 61599”; “Type” // Innsbruck / 
2/6/1914 /J. Felkel leg.” // “subsp. oenipontanus / Stauder 
/ Cotype” // “Summing / G. Warnecke / Eing. Nr. 5, 1949” 
// “ZMH 61600”; “Type” // “Innsbruck / 23.6.1913 / J. 
Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / 
G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61601”. 

Type locality. Austria: “Silltal von Wilten bei Inns- 
bruck bis Patsch, Tirol” [Sill Valley of Wilten near Inns- 
bruck to Patsch, Tirol]. 


11. serpentinicus Mayer, 1925 


Original combination. “Parn. apollo vy. serpentinicus, 
(Rbl. i. 1.) Mayer, subspecies nova” Z. des Oster. Ent.- 
Ver. 10: 2-4. 


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198 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current combination. Parnassius apollo serpentini- 
cus Mayer, 1925. 

Current status. Valid subspecies. 

Type material. Syntypes 24.4 (ZMH 824705-824706) 
(Fig. 11). “Cotype” // “Burgland / Rotlschlg. / Juli 24 / 
Mayer” // “Sammlung / G. Warnecke / Eing. Nr 5 , 1949” 
/1 “ZMH 824705”; “Cotype” // “Burgland / Stuben / Juli 
24 /L. Mayer” // “Serpentinicus Mayer” // “Sammlung / 
G. Warnecke / Eing. Nr 5 , 1949” // “ZMH 824706”. 

Type locality. Austria: “Auf einem Serpentingebirge 
an der ungarischen Grenze” [on a Serpentin mountain at 
the Hungarian Border]. 


12. ab. marschneri Bryk, 1911 


Original combination. “Parnassius apollo ab. mar- 
schneri” Bryk, 1911 Int. Ent. Z. 5: 161. 

Current combination. Parnassius apollo ab. mar- 
schneri Bryk, 1911. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
824719) (Fig. 12). “Nordeuropa / Stockholm / Umb. / 
coll. Dr. Gelpke” // “Slg. Mus. Altona / Eing. Nr. 6/1978” 
// “Type” //“Abgebildet / Bryk” // “Internat. / Entomolog. 
/ Zeitschrift /Guben / V. Jahrg. No. 20” // “ZMH 824719”. 

Original locality. Not indicated. Probably Stockholm 
[Sweden]. 

Remarks. Bryk (1911) proposed this name as an ab- 
erration of P. apollo Linnaeus, 1758, therefore, as stated 
by article 45.6.2 ICZN 1999); it is deemed to be an in- 
frasubspecific name (the author used “aberration”, “ab.”) 
and is hence unavailable. 


13. wenzeli Bryk, 1921 


Original combination. “Parnassius apollo v. Wenzeli” 
Bryk, 1921 Ent. Tidskr. 42: 117. 

Current combination. Parnassius apollo wenzeli 
Bryk, 1921. 

Current status. Valid subspecies. 

Type material. Synype 14 (ZMH 824721) (Fig. 13). 
“Cotype” // “Silltal / 19.5.1921 / J. Felkel” // “Coll. G. 
Warnecke / Eing. Nr. 11 1963” // “ZMH 824721”. 

Type locality. “Innsbruck (Patsch)”. 

Remarks. Bryk (1921) proposed this name as a va- 
riety of P. apollo Linnaeus, 1758. According to article 
45.6.4 (ICZN 1999), it is subspecific if first published be- 
fore 1961 and author expressly used the terms “variety” 
or “form”. 


14. gabrieli Bryk, 1934 


Original combination. “P. nomion ssp. gabrieli (subsp. 
nova)” Bryk, 1934 Parnassiana 3: 28. 


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Current combination. Parnassius nomion gabrieli 
Bryk, 1934. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 61602) (Fig. 14). 
“Co-Type / Collection / Kotzsch” // “Kukunor / Geb. 
Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Par- 
nassius / nomion / gabrieli 3 / Bryk” // “Coll. G. War- 
necke / Eing. Nr. 11 1963” // ZMH 61602”. 

Type _ locality. China: “Burchan-Buddha-Kette” 
[ Burchan-Buddha-Chain]. 


15. chingamensis Bryk & Eisner, 1932 


Original combination. “P. nomion F.d.W. chingamensis 
subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26. 

Current combination. Parnassius nomion chin- 
gamensis Bryk & Eisner, 1932. 

Current status. Valid subspecies. 

Type material. Syntypes 2¢'3' (ZMH 61604-61605) 
(Fig. 15). “Parnassius / chinganen- / sis 3 / C. B.-L / Chin- 
gan — Feb. / Ost- Mongolei / lec 7.” // “Coll. G. Warnecke / 
Eing. Nr. 11. 1963” //“ZMH 61604”; “Mandschur.sept.occ. 
/ Buchalu / Chingan mont. sept. / 1100 m. Juli” // “chingan- 
ensis’ // “Staudinger / Bang-Haas / Blasewitz” // “Public. 
Bryk & Eisner / Parnass. I, 2, 1932, 26 / als 1” // “Sam- 
mlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 616035”. 

Type locality. China: “Buchatu, Gudschal-Tal, Chin- 
gan mont. sept. 100 m.” [Buchatu, Gudschal-Valley, 
Chingan mont. sept. 100 m]. 

Remarks. This appears to be an erroneous original 
spelling, which was corrected by Eisner (1935) to “chin- 
ganensis”. However, Ackery (1973) treats this emenda- 
tion as an unjustified emendation according to ICZN arti- 
cle 33.2.3 (ICZN 1999) and not as a justified emendation. 


16. tsinlingensis Bryk & Eisner, 1932 


Original combination. “P. nomion F.d.W. tsinglingensis 
subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26, 

Current combination. Parnassius nomion tsinlin- 
gensis Bryk & Eisner, 1932. 

Current status. Valid subspecies. 

Type material. Syntype 1¢ (ZMH 61624) (Fig. 16). 
“Prov. Schensi / Tapaischan [?] / Tsinling schan / 3000 m 
20.6.” // “Co-Type / e Collection / Bang Haas” // “subsp. 
tsinglingensis” // “Public. Bryk t. Eisner / Parnass. II, 2, 
1932 / p. 26” // Sammlung / G. Warnecke / Eing. Nr.5, 
1949” // “ZMH 61624”. 

Type locality. China: “Schensi, Tapaischan, Tsinling- 
shan, 3000 m. VI.; Tsingschui, Liu-pinschan, 2000 m.” 


17. nominulus Staudinger, 1894 


Original combination. “Parnassius Nomion var. No- 
minulus” Staudinger, 1894 Dt. Ent. Z. Iris 7(2): 241. 


Evolutionary Systematics 5 2021, 193-261 199 


[zm » 61588 


thus Altona com: 
fcing NRPS 0B : 


ZMH - 61591 


Nordeuropa 
7 in 
tlng- 
coll. Dr. Gel pke 


Internat. 
Entomoleg. 
— Feitschrupt, 
Ube nN: 

I JSahrg He 


[zm H 824719 


e Me ee: 


ZMH 824706| 


1 cm 


Plate 2. 7. Parnassius apollo pumilus f. bispupillata Turati, 1918; 8. Parnssius apollo pumilus f. appendiculata Turati, 1918; 9. Par- 
nassius apollo ab. lacrimans Marschner, 1911; 10. Parnassius apollo oenipontanus Stauder & Felkel, 1921; 11. Parnassius apollo 
serpentinicus Mayer, 1925 12. Parnassius apollo ab. marschneri Bryk, 1911. a. Dorsal view, b. ventral view. 


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200 


Current combination. Parnassius nomion nominu- 
lus Staudinger, 1894. 

Current status. Valid subspecies. 

Type material. Type? 1419 (ZMH 824717-824718) 
(Fig. 17). “Irkutsk / Sibirien / A.M. Schmidt / Inh. M. 
Gottmann” // “Parn. nomion Hb. / Typus?” // “Coll. G. 
Warnecke / Eing. Nr. 11, 1963” // “ZMH 824717”; “Ken- 
tei-Geb.” // “Gebr. Dorries leg. 1893 / Erworb. e. coll. / 
Dorries sen. 1917” // “Nomion typ., [illegible]” // “Sig-G 
Warnecke / Eing. Nr 5 1949” // “Katalog Nr. / L. 604” // 
“ZMH 824718”. 

Type locality. Russia: “Ost- Sajan -Gebiet (stidwestlich 
von Irkutsk, an der Grenze der chinesischen Mongolei 
gelegen)” [East-Sajan region (southwest of Irkutsk, at the 
border of Chinese Mongolia)]. 

Remarks. Staudinger (1894) proposed this name as a 
variety of P. nomion Fischer & Waldheim, 1823. According 
to article 45.6.4 ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms “va- 
riety” or “form”. We believe that these two specimens be- 
long to ssp. nominulus Staudinger, 1894. The year on one of 
the labels (1893) supports this. The question mark on the la- 
bel (“Typus?’’) shows uncertainty about whether it is a type. 
In the original description, Staudinger says: “... bemerke 
ich, dass die kleineren Stticke aus dem Kentei-Gebirge den 
grosseren var. Nominulus fast gleichkommen.“ [...I realize 
that the smaller specimens from the Kentei-Mountains are 
very similar to the larger var. nominulus]. 


18. frivaldszkyi Bang-Haas, 1928 


Original combination. “Parnassius szechenyii frivaldszkyi 
O. B.- Haas, subsp. nov.” Bang-Haas, 1928 Ent. Z. 42: 60. 

Current combination. Parnassius  szechenyii 
frivaldszkyi Bang-Haas, 1928. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 61603) (Fig. 18). 
“Kansu occ. / Liang-tschou/ Richthofen mont. mar. /2500 m 
Juli” // “Co-Type / Haas” // “129” // “szechenyii / frivaldsz- 
ky?’ // “Mus Altona / Eing Nr 9-64” // “ZMH 61603”. 

Type locality. “China sept. occ.: Kansu, Richthofen 
mont., stidwestl. Uang-tschou”. 


19. ab. privatus Kotzsch, 1936 


Original combination. “Parnassius autocrator Avinoft 
ab. privatus” Kotzsch, 1936 Ent. Rdsch. 54: 44. 

Current combination. Parnassius autocrator ab. 
privatus Kotzsch, 1936. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 14 (ZMH 
61606) (Fig. 19). “Hindukusch / Chodja-Mahomed / 
Gerollzone / 3800— 4000 m 25.VII.-10. VIII. / leg. H. & 
E. Kotzsch” //“Co-Type /e Collection / Kotzsch” // “Mus 
Altona / Eing 9-64” // “ZMH 61606”. 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Original locality. Afghanistan: “Hindukusch, Chod- 
ja-Mahomed, Ger6llzone, 3800— 4000 m.” 

Remarks. Kotzsch (1936) proposed this name as an 
aberration of P. autocrator Avinoff, 1913. Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author used one of the terms 
“aberration”, “ab.” or “morph’) and is hence unavailable. 


20. lictor Bryk & Eisner, 1937 


Original combination. “P. or/eans Oberth. subsp. /ictor 
(subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58. 

Current combination. Parnassius orleans lictor 
Bryk & Eisner, 1937. 

Current status. Valid subspecies. 

Type material. Synypes 14112 (ZMH 61607-61608) 
(Fig. 20). “Kukunor/ Geb. Burchan Buddha / Nomohun 
Pass / 4000 m Juli” // “Parnassius / orleans / lictor 3 / 
Br. u. Ets.” // “Co-Type / e Collection / Kotzsch” // “Coll. 
G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61607”; 
“Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 
4000 m Juli” // “Parnassius / orleans / lictor 2 / Br. u. 
Eis.” // “Co-Type / e Collection / Kotzsch” // “Coll. G. 
Warnecke / Eing. Nr. 11, 1963” // “ZMH 61608”. 

Type locality. China: Kukunor, “Burchan-Buddha, 
Nomohun Pa”. 


21. buddha Bryk & Eisner, 1937 


Original combination. “P. mercurius subsp. buddha 
(subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58. 

Current combination. Parnassius jacquemontii 
buddha Bryk & Eisner, 1937. 

Current status. Valid subspecies. 

Type material. Syntypes 1412 (ZMH 61619- 
61620) (Fig. 21). “Co-Type / e Collection / Kotzsch” 
// “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 
4000 m Juli” // “Parnassius / mercurius | buddha & / Br. 
U. Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // 
“ZMH 61619”; “Kukunor/ Geb. Burchan Buddha / No- 
mohun Pass / 4000 m Juli” // “Co-Type / e Collection / 
Kotzsch” // “Parnassius / mercurius | buddha & / Br. U. 
Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 
61620”. 

Type locality. China: “Burchan-Buddha-Kette, No- 
mohun PaB, 4000 m.” 

Remarks. Parnassius mercurius Grum-Grshimailo, 
1891 is currently considered a subspecies of P. jacque- 
montii Boisduval, 1836. 


22. ab. tripicta Sheljuzhko, 1928 


Original combination. “Parnassius bremeri amgunen- 
sis’ 7. ab. tripicta (nov.)” Sheljuzhko, 1928 Mitt. Minch. 
Ent.Ges eis. 7; 


Evolutionary Systematics 5 2021, 193-261 201 


Kukuner 
Geb.Burchan Buddha 
Pass 


CoH. 6, Warnecke 
Eing. Nr. 17 1963 


zm : 61624 


oy “Kansu eco. : 


nahi 
ic! fen mont, mar. 
2500 mull 


Gebr. Dérries leg.1873 
Erworb. e. coll. 
DGrries sen. 1917 _ 


“Mus Altona 
sepa gente 


‘Katalog Nr. | 
CON 


ZMH 824718) 


Plate 3. 13. Parnassius apollo wenzeli Bryk, 1921; 14. Parnassius nomion gabrieli Bryk, 1934; 15. Parnassius nomion chingamen- 
sis Bryk & Eisner, 1932; 16. Parnassius nomion tsinlingensis Bryk & Eisner, 1932; 17. Parnassius nomion nominulus Staudinger, 
1894; 18. Parnassius szechenyii frivaldszkyi Bang-Haas, 1928. a. Dorsal view, b. ventral view. 


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202 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current combination. Parnassius bremeri amgun- 
ensis ab. tripicta Sheljuzhko, 1928. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 14 (ZMH 
61621) (Fig. 22). “Mandschur sept. occ. / Buchalu / Chin- 
gan mont. Sept. / 1100 m. Juli” // “Cotype” // “Co-Type 
/ e collection / Bang-Haas” // “Bremeri solonensis / ab. 
tripicta u. flavicans” // “Sammlung / G. Warnecke / Eing. 
Nr. 5, 1949” // “ZMH 61621”. 

Original locality. Not indicated. Probably “China, 
Chingan mont sept., Buchalu.” 

Remarks. Sheljuzhko (1928) proposed this name as 
an aberration of P. b. amgunensis Sheljuzhko, 1928. 
Therefore, as stated by articles 45.6.1 and 45.6.2 
(ICZN 1999); it is deemed to be an infrasubspecific 
name (the author used “aberration”, “ab.”, or the 
author expressly gave it infrasubspecific rank) and 1s 
hence unavailable. 


23. melaniticus Bang-Haas, 1915 


Original combination. “Parnassius actius melaniticus 
O. B.-H., nov. var.,” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 
7A 2A. 

Current combination. Parnassius actius melaniticus 
Bang-Haas, 1915. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 833442) (Fig. 23). 
“Co. Type” // “3 P. actius /melaniticus” //“Kutscha mont. 
/ Thianschan or / Juli 3500 m.” // “Coll. G. Warnecke / 
Eing. Nr. 11, 1963” // “ZMH 833442”. 

Type locality. China: “Tianschan or. mer., Kour- 
gak Taon, Kuldscha Gebirge, nordwestlicher Kara- 
schahr”. 

Remarks. Bang-Haas (1915) proposed this name as 
a variety of P. actius (Eversmann, 1843). According to 
article 45.6.4 (ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms 
“variety” or “form”. 


24. wernickei Kotzsch, 1936 


Original combination. “Parnassius charltonius Gray 
subsp. wernickei subsp. nova.” Kotzsch, 1936 Parnassi- 
ana 4: 6. 

Current combination. Parnassius charltonius wer- 
nickei Kotzsch, 1936. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 61622) (Fig. 24). 
“Nord-Ost-Hindukusch / Nuksan-Pass Nordseite / Al- 
penwiesenzone / 3500-3800 m Mitte Juli / leg. H. & 
E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // 
“983” // “Sig. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 
61622”, 

Type locality. Afghanistan: “Ost-Hindukusch: Nuk- 
san-PaB, Alpenwiesenzone 3500 bis 4000 m.”. 


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25. inopinatus Kotzsch, 1940 


Original combination. “Parnassius inopinatus” Kot- 
zsch, 1940 Ent. Z. 54: 17. 

Current combination. Parnassius inopinatus Kot- 
zsch, 1940. 

Current status. Valid species. 

Type material. Syntypes 6¢'¢' (ZMH 61609-61612, 
ZMH 824997-824998) (Fig. 25). “Nordwest-Afgani- 
stan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 
m 20.IV.-15.VH. / Exp. Wernicke” // “Parnassius / inopi- 
natus | f Kotzsch” // “Co-Type / Collection / Kotzsch” // 
“Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61609”; 
“Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesen- 
zone / 2800-3000 m 20.IV.-15.VII. / Exp. Wernicke” // 
“Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / 
Eing. Nr. 11, 1963” // “ZMH 61610”; “Nordwest-A fgani- 
stan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 
20.1V.-15. VII. / Exp. Wernicke” // “Co-Type / Collection 
/ Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // 
“ZMH 61611“; “Nordwest-Afganistan / Firuskuhi-Mont 
/ Alpenwiesenzone / 2800-3000 m 20.IV.-15.VII. / Exp. 
Wernicke” // “Parnassius / inopinatus /m Kotzsch” //“Co- 
Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. 
Nr. 11, 1963” // “ZMH 61612”; “Nordwest-Afghanistan / 
Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.IV.- 
15. VII. / Exp. Wernicke” // “Co-Type / e Collection / Kot- 
zsch” // “981” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // 
“ZMH 824997”; “Nordwest-A fghanistan / Firuskuhi-Mont 
/ Alpenwiesenzone / 2800-3000 m 20.IV.-15.VII. / Exp. 
Wernicke” // “Co-Type / e Collection / Kotzsch” // “980” 
//“Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824998”. 

Type locality. “Nordwest-Afghanistan, Firuskuhi-Kette, 
2800-3000 m.”. 


26. gigas Kotzsch, 1932 


Original combination. “Parnassius imperator gigas ssp. 
n.” Kotzsch, 1932 Ent. Z. 45: 267. 

Current combination. Parnassius imperator gigas 
Kotzsch, 1932. 

Current status. Valid subspecies. 

Type material. Syntypes 2¢'3' (ZMH 61613-61614) 
(Fig. 26). “Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” 
// “Parnassius / imperator / gigas f / Her.” // “Cotype” // 
“Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61613”; 
“Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” // “Par- 
nassius / imperator / gigas m / Her.” // “Cotype” // “Coll. 
G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61614”. 

Type locality. China: “vom Ostabhange des Siking- 
shan Gebirges — Sincheng etwa 2800 m.” [Eastern slopes 
of the Sikingshan Mountains — Sincheng, ca. 2800 m]. 


27. clorinda Kolar & Querci, 1937 


Original combination. “Parnassius mnemosyne L. V. 
clorinda” Kolar & Querci, 1937 Lambillionea 5: 99. 


Evolutionary Systematics 5 2021, 193-261 203 


19a 


2 


Pa 61606| 


€. Warnecke 
Ne 1, 


eeeererreraenreree: 


[zm - 61607 


gz 


Mandschur sept.oce. 
Buchalo — 


nan 
000 m 


v i 
ur 


“i 


Eing, Ne 19 


Sig. Mus, Altona 
Bing.Nr,6/1976| 


1cm 


Plate 4. 19. Parnassius autocrator ab. privatus Kotzsch, 1936; 20. Parnassius orleans lictor Bryk & Eisner, 1937; 21. Parnassius 
Jacquemontii buddha Bryk & Eisner, 1937; 22. Parnassius bremeri amgunensis ab. tripicta Sheljuzhko, 1928; 23. Parnassius actius 
melaniticus Bang-Haas, 1915; 24. Parnassius charltonius wernickei Kotzsch, 1936. a. Dorsal view, b. ventral view. 


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204 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current combination. Parnassius mnemosyne clo- 
rinda Kolar & Querci, 1937. 

Current status. Valid subspecies. 

Type material. Syntypes 2¢'¢ (ZMH 824701-824702) 
(Fig. 27). “Type” // “Macedonia / Olympus, 4500’ / June 
16, 1935 / Rome” // “Balkan / coll. Dr. Gelpke” // “Mus 
Altona / Eing. Nr. 9 — 64” // “ZMH 824701”; “Typen” // 
“Macedonia / Olympus, 4500’ / June 19, 1935 / Romei” 
// “Balkan / coll. Dr. Gelpke” // “v. clorinda Querci” // 
“P. Mnemosyne / - Clorinda Querci / Type abgebildet / in 
Lambillionea” // “Type” // “Mus Altona / Eing Nr 9-64” 
/1 “ZMH 824702”. 

Type locality. Macedonia: Olympus. 

Remarks. Kolar and Querci (1937) proposed this name 
as a variety of P. mnemosyne L. According to article 45.6.4 
(ICZN 1999), it is subspecific if first published before 1961 
and author expressly used the terms “variety” or “form”. 


28. rjabovi Sheljuzhko, 1935 


Original combination. “Parnassius mnemosyne rjabo- 
vi” Sheljuzhko, 1935 Zs. Ost. Ent. Ver. 20: 22. 

Current combination. Parnassius mnemosyne rja- 
bovi Sheljuzhko, 1935. 

Current status. Junior subjective synonym of Par- 
nassius mnemosyne caucasia Verity, 1911. 

Original material. Labelled as “Autotypes” 1319 
(ZMH 824703-824704) (Fig. 28). “autotypus 9 / rja- 
bovi Shel.” // “Kleinasien / Armenia / Daratshitshag 
/ 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. Gelpke” // 
“Mus Altona / Eing Nr 9-64” // “ZMH 824703”; “auto- 
typus 3 / rjabovi Shel.” // “Kleinasien / Armenia / Da- 
ratshitshag / 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. 
Gelpke” // “v. rjabovi Shelj.” // “Mus Altona / Eing Nr 
9-64” // “ZMH 824704”. 

Original locality. Armenia: Daratshitshag. 

Remarks. The year on the specimen labels (1939) in- 
dicate specimens were collected four years after descrip- 
tion (1935) from the same location (i.e., Armenia: Darat- 
shitshag) the original holotype material was collected. As 
a result, the label of the specimens treated as “autotypus” 
by Warnecke 1s erroneous. Autotypes do not exist in tax- 
onomy. The specimen collected from the same geograph- 
ical location from which the type specimen was collected 
is called topotype. According to ICZN (1999), a topotype 
has no formal standing and is not regulated by the Code. 
Therefore, these two specimens are erroneously labelled 
as types, should be annotated as “Not a Type” and are 
treated here as “Non-type” specimens. 


29. comitis Bryk, 1912 


Original combination. “Parnassius mnemosyne vat. co- 
mitis” Bryk, 1912 Soc. Ent. 27: 39. 

Current combination. Parnassius mnemosyne comi- 
tis Bryk, 1912. 


evolsyst.pensoft.net 


Current status. Junior subjective synonym of nomi- 
notypical Parnassius mnemosyne (Linnaeus, 1758). 

Original material. Labelled as “Cotypen” 14 (ZMH 
824707) (Fig. 29). “817” // “Italia central. / Majella / 
21.VII.1925 / coll. F Dannehl” // “Italien / coll. Dr. Gelp- 
ke” // “Cotypen” // “v. majellensis Dhl.” // “v. comitis 
Bryk.” // “Mus. Altona comm. / Eing. Nr. 3 65” // “ZMH 
824707”. 

Original locality. Italy: “aus der Majella Gruppe”. 

Remarks. Bryk (1912) proposed this name as a vari- 
ety of P. mnemosyne (Linnaeus, 1758). According to arti- 
cle 45.6.4 (ICZN 1999), it is subspecific if first published 
before 1961 and author expressly used the terms “vari- 
ety” or “form”. The year on the specimen label (1925) in- 
dicates specimen was collected 13 years after description 
(1912) from the same location (i.e., Majella Mountains, 
Italy) the original holotype material was collected. As a 
result, the label of the specimen treated as “Cotypen” is 
erroneous. The specimen collected from the same geo- 
graphical location from which the type specimen was col- 
lected is called topotype. According to ICZN (1999), a to- 
potype has no formal standing and 1s not regulated by the 
Code. Therefore, this specimen is erroneously labelled as 
type, should be annotated as “Not a Type” and is treated 
here as “Non-type” specimen. 


30. gigantea Staudinger, 1886 


Original combination. “Parn/[assius] mnemosyne L. vat. 
gigantea” Staudinger, 1886 Stett. Ent. Ztg. 47 (4-6): 197. 

Current combination. Parnassius mnemosyne gi- 
gantea Staudinger, 1886. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 824713) (Fig. 30). 
“vy. gigantea / 1/86 3 / Tary. / Ob. Turk. / “Warnecke” 
// “Tary Ob. Turkestan” // “<4 Parn. Mnemosyne | v. gi- 
gantea / E.W. BETH / Hamburg. den Haag” // “Coll. H. 
Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L / 6133” // 
“ZMH 824713”. 

Type locality. Russian Turkestan. 

Remarks. Staudinger (1886) proposed this name as a 
variety of P. mnemosyne (Linnaeus, 1758). According to 
article 45.6.4 (ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms 
“variety” or “form”. 


31. beicki Bryk & Eisner, 1932 


Original combination. “P/arnassius] epaphus beicki 
subsp. nova (O. Bang-Haas 1. |.)” Bryk & Eisner, 1932 
Parnassiana 2: 25, 42. 

Current combination. Parnassius epaphus beicki 
Bryk & Eisner, 1932. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 824714) (Fig. 31). 
“Kansu sept occ / Hsining / Nanchan mont. / Tatung / 


Evolutionary Systematics 5 2021, 193-261 205 


ZMH - 61614 


ie Ris 
ne 
~ ) 


ee 


j 
a. ey 
| Sue 


ZMH 824713 


4 
} 

\ 

4 

4 


1 cm 


Plate 5. 25. Parnassius inopinatus Kotzsch, 1940; 26. Parnassius imperator gigas Kotzsch, 1932; 27. Parnassius mnemosyne clo- 
rinda Kolar & Querci, 1937; 28. Parnassius mnemosyne rjabovi Sheljuzhko, 1935; 29. Parnassius mnemosyne comitis Bryk, 1912; 
30. Parnassius mnemosyne gigantea Staudinger, 1886. a. Dorsal view, b. ventral view. 


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206 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Nadelholzzone / Jiili 2000 m.” // “Co-Type / e Collection 
Bang-Haas” // “Ep. subsp. Beiki” // “public. v. Bryk Ut. 
Eisner / Parnass. II, 2, 1932 p. 25” // “Sammlung / G. 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 824714”. 
Type locality. China: “Std Tatung‘sche Berge, Kansu, 
3000 m* [Southern Tatung Mountains, Kansu, 3000 m]. 
Remarks. Also misspelled beiki in literature. 


32. ab. rubropicta Bang-Haas, 1915 


Original combination. “Parnassius epaphus ab. ru- 
bropicta” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 95. 

Current combination. Parnassius epaphus ab. ru- 
bropicta Bang-Haas, 1915. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Co-Type” 14 (ZMH 
824715) (Fig. 32). “Karakorum mont. / Schahidulla / 
Chines Tatarei / 4000 m. Ende Juni” // “Co-Type / e Col- 
lection / Bang-Haas” // “subtilis / rubropicata” // “S\g. 
Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824715”. 

Original locality. Not indicated. Probably China. 

Remarks. Bang-Haas (1915) proposed this name as 
an aberration of P. epaphus Oberthtr, 1879. Therefore, as 
stated by article 45.6.2 (ICZN 1999); it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


33. candidatus Bang-Haas, 1915 


Original combination. “Parnassius delphius var. candi- 
datus” Bang-Haas, 1915 Dt. Ent. Z. Iris 29 (2/3): 158. 

Current combination. Parnassius delphius candida- 
tus Bang-Haas, 1915. 

Current status. Junior subjective synonym of the 
nominotypical Parnassius delphius Eversmann, 1843. 

Type material. Syntype 14 (ZMH 824716) (Fig. 33). 
“Chin. Turkestan / Thianschan mont. or. / Juldus Tal / 
2500 m. Juli” // “Co-Type / O. Bang-Haas” // “delphius 
— caudi- | datus Cotype” // “Slg. Mus. Altona / Eing. nr. 
6/1978” // “ZMH 824716”. 

Type locality. “Tianschan or.; Juldus (Kuldsca, Ill, 
Issykkul)”. 

Remarks. Bang-Haas (1915) proposed this name as 
a variety of P. delphius Eversmann, 1843. According to 
article 45.6.4 (ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms 
“variety” or “form”. 


34. eva Kotzsch, 1936 


Original combination. “Parnassius delphius Eversm. 
subsp. eva subsp. nova” Kotzsch, 1936 Parnassiana 4: 5. 
Current combination. Parnassius jacobsoni eva 
Kotzsch, 1936. 
Current status. Valid subspecies. 


evolsyst.pensoft.net 


Type material. Syntypes 2¢¢ (ZMH 824723- 
824724) (Fig. 34). “Hindukusch / Chodja-Mahomed 
/ Gerollzone / 3800.4000 m 25.VII.-10.VIII. / leg. H. 
& E. Kotzsch” // “Co-Type / e Collection Kotzsch” // 
“O77” // “Sig. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 
824723”: “Hindukusch / Chodja-Mahomed / Gerollzone 
/ 3800.4000 m 25. VII.-10. VHT. / leg. H. & E. Kotzsch” // 
“Co-Type / e Collection Kotzsch” // “978” // “Slg. Mus. 
Altona / Eing. Nr. 6/1978” // “ZMH 824724”. 

Type locality. “Hindukusch, Chodja-Mahomed” [Af- 
ghanistan, Hindukusch, Khoja Mohammad Mts]. 


Unpublished, unavailable names 


35. ab. minor Frank 


Original combination. Parnassius mnemosyne hart- 
manni ab. minor Frank. 

Current status. “in litteris” name and hence not available. 

Original material. Labelled as “Type” 1¢ (ZMH 
824708) (Fig. 35). “Nordl. Kalkalpen / Inntaler Berge / 
Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // 
“Mus. Altona comm. / Eing. Nr. 3-65” // “Type” // “ab. 
minor Frank” // “Mus. Altona comm. / Eing Nr. 3 65” // 
“ZMH 824708”. 

Original locality. Nordl. Kalkalpen [Austria]. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. The current 
name of the subspecies is Parnassius mnemosyne hart- 
manni Standfuss, 1888. 


36. ab. nigra Frank 


Original combination. Parnassius mnemosyne hart- 
manni ab. nigra Frank. 

Current status. “in litteris” name and hence not available. 

Original material. Labelled as “Type” 1¢ (ZMH 
824709) (Fig. 36). “ab. nigra Frank” //“Nordl. Kalkalpen 
/ Chiemgau / Lofer/ 28.V.1920 / coll. Dr. Gelpke” // 
“Type” // “ab. nigra Frank” // “Mus. Altona comm. / Eing 
Nr 3 - 65” // “ZMH 824709”. 

Original locality. Nordl. Kalkalpen [Austria]. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. The current 
name of the subspecies is Parnassius mnemosyne hart- 
manni Standfuss, 1888. 


37. ab. punctata Frank 


Original combination. Parnassius mnemosyne hart- 
manni ab. punctata Frank. 


Evolutionary Systematics 5 2021, 193-261 207 


i Slg.Mus.Altona 
Eing.Nr.6/1978 
ZMH 824715 


Slg-Muse Altona 
Eing.Nr.6/1978 


ZMH 824716 


Se ee a 


wus sre ommm [MMH 824708! 


ry 


f 
sa 


2MH 824712 


Plate 6. 31. Parnassius epaphus beicki Bryk & Eisner, 1932; 32. Parnassius epaphus ab. rubropicta Bang-Haas, 1915; 33. Parnas- 
sius delphius candidatus Bang-Haas, 1915; 34. Parnassius jacobsoni eva Kotzsch, 1936; 35. Parnassius mnemosyne hartmanni 
ab. minor Frank; 36. Parnassius mnemosyne hartmanni ab. nigra Frank; 37. Parnassius mnemosyne hartmanni ab. punctata Frank; 
38. Parnassius mnemosyne hartmanni ab. nigerrima Frank. a. Dorsal view, b. ventral view. 


evolsyst.pensoft.net 


208 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current status. “in litteris” name and hence not available. 

Original material. Labelled as “Type” 1’, “Co- 
type” 14 (ZMH 824710-824711) (Fig. 37). “Cotypen” 
// “Nordl. Kalkalpen / Inntaler Berge / Oberaudorf / 
1.VI.24 500-700 m / coll. Dr. Gelpke” // “Mus. Alto- 
na comm. / Eing. Nr. 3-65” // “ZMH 824710”; “Typen” 
// “Mus. Altona comm. / Eing. Nr. 3-65” // “Nordl. 
Kalkalpen / Inntaler Berge / Oberaudorf / 30.V.23 500- 
700 m / coll. Dr. Gelpke” // “ab. punctate Frank” // 
“ZMH 824711”. 

Original locality. Nordl. Kalkalpen [Austria]. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. The current 
name of the subspecies is Parnassius mnemosyne hart- 
manni Standfuss, 1888. 


38. ab. nigerrima Frank 


Original combination. Parnassius mnemosyne hart- 
manni ab. nigerrima Frank. 

Current status. “in litteris” name and hence not 
available. 

Original material. Labelled as “Type” 14 (ZMH 
824712) (Fig. 38). “Nordl. Kalkalpen / Inntaler Berge / 
Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // 
“Type” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ab. 
nigerrima Frank” // “ZMH 824712”. 

Original locality. Nordl. Kalkalpen [Austria]. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. The current 
name of the subspecies is Parnassius mnemosyne hart- 
manni Standfuss, 1888. 


39. ab. subtusalbipupillata 


Original combination. Parnassius apollo ab. subtusal- 
bipupillata. 

Current status. “in litteris” name and hence not 
available. 

Original material. Labelled as “Type” 1¢ (ZMH 
824720) (Fig. 39). “Balkan / Bulgaria / Sofia / Mt. Vi- 
tosch / 2000 m Jul” // “ab. quincunx Bryk / ab. reniformis 
O. B.-H. / ab. laticincta Trti.” // “Mus. Altona comm. / 
Eing. Nr. 3-65” // “Type” // ZMH 824720”. 

Original locality. Balkan, Bulgaria. 

Remarks. This taxon cannot be found in the 
literature. As stated by article 45.6.2 (ICZN 1999); it 
is deemed to be an infrasubspecific name (the author 
used “aberration”, “ab.”) and is hence unavailable. 
The current name of the species 1s Parnassius apollo 
(Linnaeus, 1758). 


evolsyst.pensoft.net 


40. ab. analifasciata Gelpke 


Original combination. Parnassius apollo ab. analifas- 
ciata Gelpke. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Type” 1¢ (ZMH 
824722) (Fig. 40). “Stiddeutschland / Frank. Jura / Eich- 
statt / 12.VII.1921 / coll. Dr. Gelpke” // “Type” // “ab. 
analifasciata Gelpke” // “ab. splendida Osth. —fasciata st. 
/ ab. magnifica Bryk / ab. subt-bip. Fr. ab. subt-graphica / 
ab. seitzi Brk. — embryki Bryk.” // “Mus. Altona comm. / 
Eing. Nr. 3-65” // “ZMH 824722”. 

Original locality. South Germany. 

Remarks. This taxon cannot be found in the literature. 
As stated by article 45.6.2 (ICZN 1999); it is deemed to 
be an infrasubspecific name (the author used “aberra- 
tion”, “ab.”) and is hence unavailable. The current name 
of the species is Parnassius apollo (Linnaeus, 1758). 


Subfamily Papilioninae Latreille, 1802 
Tribe Papilionini Latreille, 1802 


41. f. unimaculata Warnecke, 1924 


Original combination. “Papilio xuthus Linnaeus f. uni- 
maculata” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149. 

Current combination. Papilio xuthus f. unimacula- 
ta Warnecke, 1924. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 1¢ (ZMH 
61623) (Fig. 41). “Japan” // “Type” // Sammlung / Fritz 
Diehl / Eing. Nr. 35, 1950” // “P. xuthus L. / n.f. // uni- 
maculata / Warn.” // “ZMH 61623”. 

Original locality. Original publication without loca- 
tion records; label states Japan; according to hand written 
type catalogue of Warnecke: Ussuri ex Coll. Dorries (67), 
holotype destroyed in fires in Zool. Mus. Hamburg 1943, 
1 specimen with location label Japan labelled as type by 
Warnecke in Coll. Warnecke, now in ZMH. 

Remarks. Warnecke (1924) proposed this name as a 
form of P. xuthus Linnaeus, 1776. According to article 
45.6.1 (ICZN 1999), it is infrasubspecific if the content 
of the work unambiguously reveals that the name was 
proposed for an infrasubspecific entity, and 1s hence un- 
available. The text of Warnecke (1942) paper unambigu- 
ously reveals that he describes an individual form and not 
a (geographic) subspecies. 


Tribe Troidini Talbot, 1939 


42. peruanus Fuchs, 1954 


Original combination. “Papilio archidamas var. peruanus” 
Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 83-84. 
Current combination. Battus polydamas peruanus 
(Fuchs, 1954). 
Current status. Valid subspecies. 


Evolutionary Systematics 5 2021, 193-261 


Type material. Lectotype 14 (ZMH 61615) (Fig. 42) 
Paralectotypes 2312 (ZMH 61616-61617, ZMH 
824999). “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 
2000 m / Type” // “HOLOTYPE / Papilio archidamas / 
var. peruanus / von Fuchs” // “archidamas / S. Rio Pam- 
pas / 2000 m” // “Z.1.M. / Hamburg” // “LECTOTYPE 3 
/ Papilio archidamas / var. peruanus Fuchs / by G. La- 
mas, 1978” // “Coll. H. Rédinger / Eing. Nr. 30, 1952 
/ Katalog Nr. L.” // “Battus archidamas / peruanus von 
Fuchs / G. Lamas. Det. 1978” // “Battus polydamas / pe- 
ruanus (Fuchs, 1954) / Lectotype 3 / Racheli & Pisched- 
da 1987” // “ZMH 61615”; “Rio Pampas / Cuesta de 
Bonbon / 8/1928 / ca. 2000 m / Type” // “P. archidamas 
peruanus” // “Z.1.M. Hamburg” // “Battus polydamas / 
peruanus (Fuchs, 1954) / Paralectotype 3 / Racheli & 
Pischedda, 1987” // “ZMH 61616”; “Rio Pampas / Cues- 
ta de Bonbon / 8/1928 / ca. 2000 m / Type” // “Battus 
polydamas | peruanus (Fuchs, 1954) / Paralectotype 9 
/ Racheli & Pischedda, 1987” // “Z.1.M. Hamburg” // 
“ZMH 61617”; “3” // “Papilio / archidamas & / var. pe- 
ruanus / von Fuchs / Rio Pampas / Cuesta de Bonbon / c. 
2000 m / 8/28” // “Z.I.M. / Hamburg” // “Type” // “Battus 
polydamas / peruanus (Fuchs, 1954) / Paralectotype 3 / 
Racheli & Pischedda 1987” // “Coll. H. Rodinger / Eing. 
Nr. 30, 1952 / Katalog Nr. L.” // “ZMH 824999”. 

Type locality. [Lectotype] Peru: “Rio Pampas, Cues- 
ta de Bonbon, ca. 2000 m”; lectotype designated by La- 
mas (1978). 

Remarks. Fuchs (1954) proposed this name as a va- 
riety of P. archidamas Boisduval, 1836. According to 
article 45.6.4 (ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms 
“variety” or “form”. Papilio archidamas Boisduval, 1836 
is a junior subjective synonym of Battus polydamas (Lin- 
naeus, 1758). 


Tribe Leptocircini Kirby, 1896 
43. ab. ornatissima Warnecke, 1924 


Original combination. “Papilio podalirius L. n. ab. or- 
natissima Warn.” Warnecke, 1924 Int. Ent. Z. 18: 157. 

Current combination. [phiclides podalirius ab. or- 
natissima (Warnecke, 1924). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
61618) (Fig. 43). “n. ab. ornatissima / Warnecke. Type” // 
“Type” // “altester [?] Name: confluens Ver. / Rhop. Pal. 
P. 292, t. 57, f. 5, 1911 / vgl Lambillioneaa 1932 p. 223” // 
“Katalog Nr. / L. 204” // “Slg. G. Warnecke / eing. Nr. 5. 
1949” // “ZMH 61618”. 

Original locality. Southern Germany. 

Remarks. Warnecke, (1924) proposed this name as an 
aberration of Papilio podalirius Linnaeus, 1758. There- 
fore, as stated by article 45.6.2 (ICZN 1999) it is deemed 
to be an infrasubspecific name (the author used one of 
the terms “aberration”, “ab.” or “morph”) and is hence 
unavailable. 


209 
Unpublished, unavailable names 


44. ab. indistinctus Meinhard 


Original combination. [phiclides feisthamelii f. ornat- 
us ab. indistinctus Meinhard. 

Current status. “in litteris” name and hence not available. 

Original material. Labelled as “Type” 14 (ZMH 
835574) (Fig. 44). “26/5. 1906 / Stuttgart / ex larv.” // 
“ab. indistinctus / Meinh.” // “ZMH 835574”. 

Remarks. This taxon cannot be found in the literature. 
Probably an aberration of Jphiclides feisthamelii (Du- 
ponchel, 1832). Apparently the imago emerged in Stutt- 
gart (Germany), but the larva must have been collected in 
the Iberian Peninsula (M. Wiemers pers.com. ). 


Family Pieridae Swainson, 1820 
Subfamily Coliadinae Swainson, 1827 


45. ab. crocopepla Warnecke, 1912 


Original combination. “Colias aurora Esp. 9 ab. croco- 
pepla” Warnecke, 1912 Soc. Ent. 27: 68. 

Current combination. Colias heos ab. crocopepla 
Warnecke, 1912. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type’12 (ZMH 
827594) (Fig. 45). “Type” // “Bikienfl.” // “ab. crocope- 
pla / Warn. Soc. Ent. 27 6. 68” // “L.6” // “sig. G. War- 
necke / Eing. Nr. 5, 1949” // “Katalog Nr. / L 1754” // 
“AMUR-GEBIET: / Gebr. Dorries leg. 1887 / Erworb. e. 
coll. / Doérries sen. 1917 / (vgl. Mém. Rom. VI, / 1892 p 
91)” // “ZMH827594”. 

Original locality. Amur [Russia]. 

Remarks. Papilio aurora Esper, 1783 is a junior pri- 
mary homonym of Papilio aurora Cramer, [1780]. The 
former is currently known as Colias heos (Herbst, 1792) 
and the latter is known as Colotis aurora (Cramer, [1780]). 
Grieshuber and Lamas (2007) provided a nomenclatural 
list of all available and unavailable names in the genus 
Colias and clearly stated that Colias aurora [ab.| croco- 
pepla is infrasubspecific and thus not available name. 


46. ab. rhododactyla Warnecke, 1912 


Original combination. “Colias aurora Esp. 9 ab. rhodo- 
dactyla” Warnecke, 1912 Soc. Ent. 27: 68. 

Current combination. Colias heos ab. rhododactyla 
Warnecke, 1912. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 192 (ZMH 
827593) (Fig. 46). “L.39 / Col. aurora/ ab. 2 rhododactyla / 
Warn.” // “ab. rhododactyla / Warnecke / Soc. Ent. 27, 1912 
p. 68” //“Katalog Nr. /L. 1755” //“Tjutyuché / Ost-Siberien” 
// “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827593”. 

Original locality. Amur [Russia]. 


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210 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Remarks. Papilio aurora Esper, 1783 is a junior pri- 
mary homonym of Papilio aurora Cramer, [1780]. The 
former is currently known as Colias heos (Herbst, 1792) 
and the latter is known as Colotis aurora (Cramer, [1780]). 
Grieshuber and Lamas (2007) provided a nomenclatural 
list of all available and unavailable names in the genus 
Colias and clearly stated that Colias aurora [ab.| rhodo- 
dactyla is infrasubspecific and thus not available name. 


47. ab. theia Warnecke, 1912 


Original combination. “Colias aurora Esp. & ab. theia” 
Warnecke, 1912 Soc. Ent. 27: 68. 

Current combination. Colias heos ab. theia War- 
necke, 1912. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Lectotype” 12° (ZMH 
827609) (Fig. 47). “Kentei- / Gebirge” // “Gebr. Dorries 
leg.1889 / Erworb. E. coll. / Dorries sen. 1917” // “f. theia 
Warnecke / Die Type ist im Zool. Mus. Hamburg!” // “L. 
38 / Lekto-Type / Col. aurora f. theia warn.” // “Katalog 
Nr. /L 1750” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // 
“ZMH 827609”. 

Original locality. Amur [Russia]. 

Remarks. Papilio aurora Esper, 1783 is a junior pri- 
mary homonym of Papilio aurora Cramer, [1780]. The 
former is currently known as Colias heos (Herbst, 1792) 
and the latter is known as Colotis aurora (Cramer, [1780]). 
Grieshuber and Lamas (2007) provided a nomenclatural 
list of all available and unavailable names in the genus 
Colias and clearly stated that Colias aurora [ab.] theia is 
infrasubspecific and thus not an available name. 


48. f. rebeli Schawerda, 1906 


Original combination. “Colias myrmidone balcdnica f. 
rebeli” Schawerda, 1906; Verh. Zool. Bot. Ges. Wien 6: 
651. 

Current combination. Colias caucasica balcanica f. 
rebeli Schawerda, 1906. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 12 (ZMH 
827619) (Fig. 48). “Orjen / Herzegowina / Dr. Schaw- 
erda” // “Sammlung / Fritz Diehl / Eing. Nr. 35,1950” // 
“rebeli 2” // “Cotype” // “Cotype” // “LS” // “Katalog Nr. 
L 1721” // “ZMH 827619”. 

Original locality. Bosnia and Herzegovina: 
Trebevic”’. 

Remarks. Schawerda (1906) proposed this name as a 
form of C. m. balcanica Rebel, 1901. Therefore, as stat- 
ed by article 45.6.1 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author expressly gave it infra- 
subspecific rank) and 1s hence unavailable. Colias bal- 
canica Rebel, 1901 is currently regarded as subspecies 


(<4 


vom 


evolsyst.pensoft.net 


of C. caucasica Staudinger, 1871. Grieshuber and Lamas 
(2007) provided a nomenclatural list of all available and 
unavailable names in the genus Colias and clearly stat- 
ed that Colias myrmidone balcanica |f.| rebeli is infra- 
subspecific and thus not available name. 


49. tatarica Bang-Haas, 1915 


Original combination. “Colias cocandica tatarica O. B.- 
H. n. var.” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 98. 

Current combination. Colias cocandica tatarica 
Bang-Haas, 1915. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 833446) (Fig. 49). 
“Co-Type” // “Chotan mer / Karakorum / Juli” // “VII. 
/ Chotan. mer. / Karakorum / v. Friedrich [?]” // “ZMH 
833446”. 

Type locality. China: “... am Nordabhange des Kara- 
korum, in den Gebirgen stidlich der Stadt Chotan (Chi- 
nesische Tatarei)” [...around the northern slopes of the 
Karakorum, in the Mountains south of the town of Cho- 
tan (Chinese Tartare) ]. 

Remarks. Bang-Haas (1915) proposed this name as 
a variety of C. cocandica Erschoff, 1874. According to 
article 45.6.4 (ICZN 1999), it is subspecific if first pub- 
lished before 1961 and author expressly used the terms 
“variety” or “form”. 


50. aurea Kotzsch, 1936 


Original combination. “Colias wiskotti aurea subsp. n.” 
Kotzsch, 1936 Ent. Rdsch. 54(4): 45. 

Current combination. Colias wiskotti aurea Kot- 
zsch, 1936. 

Current status. Valid subspecies. 

Type material. Syntypes 3¢¢ 19 (ZMH 827622- 
827623, ZMH 835580—-835581) (Fig. 50). “Nord-Ost- 
Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone 
/ 3500-4000 m Mitte Juli/leg. H. & E. Kotzsch” // “Wisk. 
aurea Kotzsch” // “Co-Type / e Collection / Kotzsch” // 
“L.7” //“Katalog Nr. /L 1793” //“Sig G. Warnecke / Eing. 
Nr. 5, 1946” // “ZMH 827622”; “Nord-Ost-Hindukusch / 
Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500—4000 
m Mitte Juli / leg. H. & E. Kotzsch” // “Katalog Nr. / L 
1794” // “Wiskotti aurea Ktz” // “Co-Type / e Collection / 
Kotzsch” //“L.8” // “Sig G. Warnecke / Eing. Nr. 5, 1946” 
// “ZMH 827623”; “Nord-Ost-Hindukusch / Nuksan- 
Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte 
Juli / leg. H. & E. Kotzsch” // “Co-Type / e Collection 
/ Kotzsch” // “Wiskotti 4/ aurea’ // “ZMH 835580”: 
“Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite  / 
Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H. & E. 
Kotzsch” // “Colias 9° / wiskotti / v. aurea” // “Co-Type / 
e Collection / Kotzsch” // “ZMH 835581”. 

Type locality. Afghanistan:“Ost-Hindukusch, Nord- 
seite, Alpenwiesenzone, 3500-3800 m.” 


Evolutionary Systematics 5 2021, 193-261 211 


Siiddeutschland 
Frank. Jura he 


Eichstatt v7 


Balkan: coll. Dr, Gelpke sgzy 
BULGARIA et | — 
2000 aul | | Pp yD e | | 
ab. quincunx rac 
4b. seniors OB. 


| abanalifasciala Gelnas 


a6. spon dicta dst fasctala. 
af sabtbsn takes repli 
ab sectry Bat emBryne be 


Mus. A tona comm | 
ting. NF, 3-64 ZMH 824722 


ab laticinclareh. 


Mus. Altona comin 
Fing Ne 3+ 65 


Type _ | 


[zm 824720 


ee 
Z.bM. 
Hamner | 


4 


by & lames, (978 


| Battus pelydseas 
| pervanvs (Fecks 1454) 
Lectotypéee? 
Rachel’ & Pischeclela (98% 


ry 


ZMH * 61615 


|ZMH 835574 


Plate 7. 39. Parnassius apollo ab. subtusalbipupillata; 40. Parnassius apollo ab. analifasciata Gelpke; 41. Papilio xuthus f. unimac- 
ulata Warnecke, 1924; 42. Battus polydamas peruanus (Fuchs, 1954); 43. Iphiclides podalirius ab. ornatissima (Warnecke, 1924); 
44. Iphiclides feisthamelii f. ornatus ab. indistinctus Meinhard. a. Dorsal view, b. ventral view. 


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Z2N2 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


ZMH 827593 


Orjen 3. 
Herzegowina 


“ 
Dr.Sghawerda 


Sammlung 
: Eritz Diehl + 
Eing.Nr95,1950 ! 


ZMH 827609 


ZMH 827619 


Nord-Ost-Hindukuseh_ 
Nuksan-Pass-Nordseite 
: enwitsenzone 

: Mitte Juli 
leg.H.&EKotzsch 


sult bs 


11cm 


Plate 8. 45. Colias heos ab. crocopepla Warnecke, 1912; 46. Colias heos ab. rhododactyla Warnecke, 1912; 47. Colias heos ab. 
theia Warnecke, 1912; 48. Colias caucasica balcanica f. rebeli Schawerda, 1906; 49. Colias cocandica tatarica Bang-Haas, 1915; 


50. Colias wiskotti aurea Kotzsch, 1936. a. Dorsal view, b. ventral view. 


evolsyst.pensoft.net 


Evolutionary Systematics 5 2021, 193-261 


51. yauta Fuchs, 1928 


Original combination. “T7eriocolias yauta’ Fuchs, 1954 
Beitrage zur Fauna Perus 4:83. 

Current combination. Eurema Zelia andina (Fuchs, 
1928). 

Current status. Junior subjective synonym of Eurema 
zelia andina (Forbes, 1928). 

Type material. Holotype 14 (ZMH 825001), Para- 
types 80:2 (ZMH 825000, ZMH 825002-825008) (Fig. 
51). “Teriocolias / yauta 3 / J. Fuchs / Cordillere von / 
Tlaula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. 
Nr. 4. 1956” // “Type” // “Paratype / Teriocolias yauta / 
von Fuchs” // “Teriocolias zelia | andina Forbes / G. La- 
mas. det. 1978” // “ZMH 825000”; “Llauta / ca. 3000 m / 
5/1922 / Type” // “Coll. H. R6dinger / Eing. Nr. 30, 1952” 
// “Holotype / Teriocolias yauta / von Fuchs” // “Terioco- 
lias zelia | andina Forbes / G. Lamas. det. 1978” // “ZMH 
825001”; “Llatita / ca. 3000 m / 5/1922 / Type” // “Coll. H. 
Rodinger / Eing. Nr. 30, 1952” // “ZMH 825002”; “Lara- 
mate / ca 3400 m / 5/1922 / Type” // “Coll. H. Roding- 
er / Eing. Nr. 30, 1952” // “ZMH 825003”; “Oberhalb / 
Aina / ca 3000 m / 8/1924 / Type” // “Coll. H. R6dinger / 
Eing. Nr. 30, 1952” // “ZMH 825004”; “Llatita / ca. 3000 
m / 5/1922 / Type” // “Coll. H. Rodinger / Eing. Nr. 30, 
1952” // “ZMH 825005”; “Type” // Teriocolias / yauta 9 / 
v. Fuchs / Cordillere von / Ilaula 5.22 / 3000 m” // “Sam- 
mlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “ZMH 825006”; 
“Type” // “Sid — Peru / Ocana, 2600 m / 7.4.1936.” // 
“Hbg. — Std — Peru / Sammelreise 1936. / Eing. Nr. 1, 
1937.” //“Teriocolias / yauta 3 / v. Fuchs” // “Sammlung / 
W, v Fuchs / Eing. Nr. 4, 1956” // “ZMH 825007”; “Type” 
/! Teriocolias | yauta 2 / v. Fuchs / Cordillere von / Taula 
5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 
1956” // “ZMH 825008”. 

Type locality. South Peru. “ ... Kordillere von Llauta 
... auch an anderen Stellen der Kordilleren” ” [Cordillera 
of Llauta...also other areas in the cordilleras]. 


Subfamily Dismorphiinae Schatz, 1887 


52. lorkovici Pfeiffer, 1932 


Original combination. “Leptidea duponcheli Stgr. ssp. 
n. Jorkovici m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 
1932, Mitt. Munch. Ent. Ges. 22: 20. 

Current combination. Leptidea duponcheli lorkovici 
Pfeiffer, 1932. 

Current status. Valid subspecies. 

Type material. Syntypes 244 (ZMH 827614827615) 
(Fig. 52). “Syr. Sept. / Taurus c. / Marasch / 6-900 m V.30 
/ Einh. Slr. leg.” // “Cotypus / duponcheli / ssp. lorkovici 
/ Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // 
“ZMH 827614”; “Syria s. / Amanus s. / (Duldtil Dagh) / 
Jeschildere / VI 33 / Coll. Pfeiffer / Mtinchen” // “Coty- 
pus / duponcheli / ssp. lorkovici / Pfeiffer” // “Coll. By- 
tinski-Salz / Eing. Nr 20 - 60” // “ZMH 827615”. 

Type locality. Turkey: “Marasch in tirkisch Nordsy- 
rien; An den Héngen um M bis 1200 m” [Marasch in the 


213 


turkish regions of northern Syria. On the slopes around 
Marasch up to 1200 m]. 


53. f. fragilis Dannehl, 1925 


Original combination. “Leptidea sinapis L. (gen. autum- 
nalis) v. fragilis Dannehl” Dannehl, 1925 Ent. Z. 39: 6. 

Current combination. Leptidea sinapis f. fragilis 
Dannehl, 1925. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 14 (ZMH 
835575) (Fig. 53). “3.10.24” // “Sudtirol Terlan / Franz 
Dannehl” // “fragilis Dhl. ” // “Cotype” // “3.10 1924 / 
Terlan / Siidtirol / Dannehl” // “ZMH 835575”. 

Original locality. Italy: “in Stdtirol bei Bozen, Fle- 
imstal, Nonstal.” 

Remarks. Dannehl (1925) proposed this name as a 
form of L. sinapis (Linnaeus, 1758). According to article 
45.6.1 ICZN 1999), it is infrasubspecific if the content of 
the work unambiguously reveals that the name was pro- 
posed for an infrasubspecific entity, and is hence unavail- 
able. The text of Dannehl (1925) paper unambiguously 
reveals that this form was described as a seasonal form 
(autumn generation) and not as a subspecies. 


54. ab. simbruina Dannehl, 1927 


Original combination. “Leptidea sinapis L. ab. simbru- 
ina Dhl.” Dannehl, 1927 Mitt. Mtinch. Ent. Ges. 17: 1. 

Current combination. Leptidea sinapis ab. simbru- 
ina Dannehl, 1927. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 2¢¢ (ZMH 
835582—835583) (Fig. 54). “25.6” // “Italia central. / Mti. 
Simbruini / coll. F. Dannehl” // “simbruina” // “Cotypen” 
/1 “25.6. 1927 / Mte Simbruini / Central Italien” // “ZMH 
835582”; “7.4. 1927 / Mte Simbruini / Central Italien” // 
“ZMH 835583”. 

Original locality. Italy: “Sabiner und Italy: Simbru- 
iner Berge vom Tal (Campagna) bis auf die Gipfel der 
angegebenen Gebirge.” [Sabiner and Italy: Simbruin 
Mountains from the valley (Campagna) up to the top of 
the stated mountain ranges]. 

Remarks. Dannehl (1927) proposed this name as an 
aberration of L. sinapis (Linnaeus, 1758). Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author used one of the terms 
“aberration”, “ab.” or “morph”) and is hence unavailable. 


55. stabiarum Staudinger, 1914 


Original combination. “Leptidea (Leucophasia) sina- 
pis stabiarum m., nov. subspec.” Staudinger, 1914 Zeit. 
Wiss. Ins. Biol. 10: 371, t. 11, figs. 5, 6. 


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214 


Current combination. Leptidea sinapis stabiarum 
Staudinger, 1914. 

Current status. Junior subjective synonym of nomi- 
notypical Leptidea sinapis (Linnaeus, 1758). 

Type material. Syntypes 3¢¢ (ZMH 827624— 
827626) (Fig. 55). “Ital. mer. 600 m. / penins. Surren- 
tina / M. Faito 3.6.1920 / H. Stauder legit.” // “Type” // 
“Sammlung / G Warnecke / Eing Nr 5, 1949” // “ZMH 
827624”; “25/6 / Ital. mer. m. / penins. Surrentina / M. 
Fatio 3.6.1920 / H. Stauder legit.” // “Type” // “Sammlu- 
ng / G Warnecke / Eing Nr 5, 1949” // “ZMH 827625”; 
26/6 / Ital. mer. 700 m. / penins. Surrentina / M. Fatio 
3.6.1920 / H. Stauder legit.” // “Type” // “Sinapis stabi- 
arum / Stauder / v. stabiarum.” // “ZMH 827626”. 

Type locality. Italy: Monte Faito and S. Angelo [Na- 
ples], 700-1400 m. 


Subfamily Pierinae Swainson, 1820 
56. italorum Dannehl, 1929 


Original combination. “Anthocharis euphenoides St. italo- 
rum Dhl.” Dannehl, 1929 Mitt. Munch. Ent. Ges. 19: 98. 

Current combination. Anthocharis euphenoides 
italorum Dannehl, 1929. 

Current status. Junior subjective synonym of 
nominotypical Anthocharis euphenoides Staudinger, 1869. 

Type material. Syntype 14 (ZMH 835584) (Fig. 56). 
“20.5” // “Apenn. Central. / Mte. Genzana / 1500—2000 
m. / coll. F. Dannehl” // “Cotype” // “20.5.1928 / Mte 
Genzana / Central- Apenninen / 1500-2000 m D.” // 
“ZMH 835584”. 

Type locality. Italy: in den Gebieten des 
Velino, Sirente, Monte Genzana, Monte Paradiso 
und der Majella. auf Hohen von 1500-2000 m.” [in 
the mountain ranges of the Velino, Sirente, Monte 
Genzana, Monte Paradiso and Majella. In altitudes of 
1500-2000 m]. 


(<4 


57. paravicinii Stauder, 1915 


Original combination. “Euchloé belia paravicinii”’ 
Stauder, 1915 Dt. Ent. Z. Iris 29: 25. 

Current combination. Euchloe ausonia paravicinii 
Stauder, 1915. 

Current status. Junior subjective synonym of Eu- 
chloe ausonia melanochloros Rober, 1907. 

Type material. Syntype 1¢ (ZMH 827616) 
(Fig. 57). “Algeria mer. / Djebel Aures / 10.5.1914 / 
H. Stauder” // “Paravicinii Stauder” // “paravicinii 
Stauder” // “Type” // “L. 4” // “Katalog Nr. / L. 1180” 
// “Type” // “Sig. G. Warnecke / Eing. Nr. 5, 1948” // 
“ZMH 827616”. 

Type locality. Algeria: “Djebel Aureés in Stidalgerien”’. 

Remarks. Papilio belia Stoll [1782] is a primary hom- 
onym of Papilio belia Linnaeus, 1767 which is currently 
known as Anthocharis belia. 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


58. f. flavide apicata Stauder, 1922 


Original combination. “7eracolus daira nouna auresia- 
ca g. II. interposita Stdr. f. flavide apicata m. nov.” Staud- 
er, 1922 Mitt. Mitinch. Ent. Ges. 12: 24—25. 

Current combination. Colotis evagore nouna f. fla- 
vide apicata (Stauder, 19272). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 74° (ZMH 
827595-827599, ZMH 827601-827602) (Fig. 58). 
“Type” // “El Kautara / auf VI” // “Auresica gen. II. / in- 
terposita Stauder” // “Sammlung / G. Warnecke / Eing. 
Nr. 5, 1949” // “ZMH 827595”: “2” // “Type” // “Algeria 
mer. / Djebel Aures / VI. 1913 / H. Stauder” // “f. inter- 
posita | 5.VY’ // “VI form / 3 200” // “Sammlung / G. 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827597”; “Type” 
// “El Kautara / auf VI’ // “Sammlung / G. Warnecke / 
Eing. Nr. 5, 1949” // “ZMH 827596”; “Type” // “El Kau- 
tara / auf VI” // “Sammlung / G. Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827598”; “3” // “Type” // “Algeria mer. / 
Djebel Aures / VI. 1912 / H. Stauder” // “Sammlung / G. 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827599”; “Type” 
//“X Dyebel Aures / VII. 1912 H. Stauder” // “Sammlung 
/ G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827601”; 
“Q” // “Type” // “Algeria mer. / Djebel Aures / VI. 1913 
/ H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827602”. 

Original locality. Not indicated. Probably “Algeria 
mer. / Djebel Aures”’. 

Remarks. Stauder (1922) proposed this name as 
a form of 7! d. nouna auresiaca interposita Staud- 
er, 1922. Therefore, as stated by article 45.6.1 (ICZN 
1999) it is deemed to be an infrasubspecific name (the 
author expressly gave it infrasubspecific rank) and 1s 
hence unavailable. Teracolus daira nouna Lucas, 1849 
is junior subjective synonym of Colotis evagore nouna 
(Lucas, 1849). 


59. f. vernalis alticola auresiaca Stauder, 1913 


Original combination. “T7eracolus daira (= dalia Fldr.) 
nouna Luc. 1848 (= demagore Fldr.) forma vernalis alti- 
cola auresiaca” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290. 

Current combination. Colotis evagore nouna f. ver- 
nalis alticola auresiaca (Stauder, 1913). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 3939 (ZMH 
827600, ZMH 827603-827607) (Fig. 59). “3” // “Co- 
type” // “Algeria mer. / Djebel Aures / V. 1912 / H. Staud- 
er” // “auresiaca” // “Sammlung / G. Warnecke / Eing. 
Nr. 5, 1949” // “ZMH 827600”; “2” // “Type” // “Algeria 
mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca / 
S Stdr.” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” 
/1 “ZMH 827603”; “9” // “Type” // “Algeria mer. / Djebel 
Aures / V. 1912 / H. Stauder” // “auresiaca Stdr. / 2” // 


Evolutionary Systematics 5 2021, 193-261 215 


Coll. H. ‘Rbdinger 
Eing. Nr, 30, 1992 


TYPE 
FO otaS gousbe ‘Goll, Bytinski-Salz: 
you Fucks teing. Ns. 20, 1960: 
Teyocokkas 2eka 
Anokne. forbes 


G. Lamas. det. 1978 ; 


ZMH 825001| 


Apenn.central. 
Mite. Génzana’ 
1500-2000 m 


hee 


nna se 


[2M 827616 


1 cm 


Plate 9. 51. Eurema zelia andina (Fuchs, 1928); 52. Leptidea duponcheli lorkovici Pfeiffer, 1932; 53. Leptidea sinapis f. fragilis 
Dannehl, 1925; 54. Leptidea sinapis ab. simbruina Dannehl, 1927; 55. Leptidea sinapis stabiarum Staudinger, 1914; 56. Antho- 
charis euphenoides italorum Dannehl, 1929; 57. Euchloe ausonia paravicinii Stauder, 1915; 58. Colotis evagore nouna f. flavide 
apicata (Stauder, 1922). a. Dorsal view, b. ventral view. 


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216 


“Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 
827604”: “9” // “Cotype” // “Algeria mer. / Djebel Aures 
/V. 1912 /H. Stauder” // “auresiaca / 9” // “Sammlung / 
G Warnecke / Eing. Nr. 5,1949” // “ZMH 827605”; “2” 
// “Cotype” // “Algeria mer. / Djebel Aures / V. 1912 / H. 
Stauder” // “auresiaca / 2” // “Sammlung / G Warnecke 
/ Eing. Nr. 5,1949” // “ZMH 827605”; “4” // “Type” // 
“Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // 
“auresiaca Stdr. / 3” // “Sammlung / G Warnecke / Eing. 
Nr. 5,1949” // “ZMH 827607”. 

Original locality. Algeria: “in den algerischen Atlas 
und die nordliche Sahara” [in the Algerian Atlas and the 
northern Sahara]. 

Remarks. Stauder (1913) proposed this name as a 
form of 7: d. nouna (Lucas, 1848). Therefore, as stated 
by article 45.6.1 ICZN 1999) it is deemed to be an in- 
frasubspecific name (the author expressly gave it infra- 
subspecific rank) and is hence unavailable. Zeracolus 
daira nouna Lucas, 1849 is junior subjective synonym of 
Colotis evagore nouna (Lucas, 1849). 


60. f. biformata Stauder, 1913 


Original combination. “T7eracolus daira (= dalia Fldr.) 
nouna Luc. 1848 (= demagore Fldr.) forma autumnalis 
biformata m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290. 

Current combination. Colotis evagore nouna f. bi- 
formata (Stauder, 1913). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827608) (Fig. 60). “4” // “biformata / < Stdr.” // “Type” // 
“Algeria mer. / Djebel Aures / V. 1912 /H. Stauder” //“Sam- 
mlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827608”. 

Original locality. Algeria: “in den algerischen Atlas 
und die nordliche Sahara” [in the Algerian Atlas and the 
northern Sahara]. 

Remarks. Stauder (1913) proposed this name as a 
form of 7! d. nouna (Lucas, 1848). Therefore, as stated 
by article 45.6.1 (ICZN 1999) it is deemed to be an in- 
frasubspecific name (the author expressly gave it infra- 
subspecific rank) and is hence unavailable. Teracolus 
daira nouna Lucas, 1849 is junior subjective synonym of 
Colotis evagore nouna (Lucas, 1849). 


61. f. aestiva pyroleuca Stauder, 1913 


Original combination. “7eracolus daira nouna Luc. for- 
ma aestiva pyroleuca m.” Stauder, 1913 Zeit. Wiss. Ins. 
Biol. 9: 290. 

Current combination. Colotis evagore nouna f. aes- 
tiva pyroleuca (Stauder, 1913). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type and Cotype” 
1049 (ZMH 827618, ZMH 827628-827636) (Fig. 61). 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


“O” // “Dyjebel Aures / VII. 1912 Stauder” // “nouna 
var. / pyroleuca / Stauder” // “TYPE” // “Sammlung / G 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827618”; “2” // 
“Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // 
“pyroleuca 2” //“COTYPE” // “Sammlung / G Warnecke 
/ Eing. Nr. 5, 1949” // “ZMH 827628”; “3” // “Algeria 
mer. / Djebel Aures / VII. 1912 / H. Stauder” // “Cotype” 
// “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827629”, “3” // “Algeria mer. / Djeb- 
el Aurés / VII. 1912 / H. Stauder” // “pyroleuca 4” // 
“TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” 
/1 “ZMH 827630”; “2” // “Algeria mer. / Djebel Aurés / 
VII. 1912 /H. Stauder” // “pyroleuca 9” // ““COTYPE” // 
“Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 
827631”: “3” // “Algeria mer. / Djebel Aurés / VII. 1912 / 
H. Stauder” // “pyroleuca 6” // “COTY PE” // “Sammlung 
/ G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827632”; “2” 
// “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” 
// “pyroleuca 3 NO” // “COTYPE” // “Sammlung / G 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827633”; “3” // 
“COTYPE” // Algeria mer. / El Kaictara [?] / anfg [IX 19 / 
H. Stauder” // “pyroleuca 4” // “Sammlung / G Warnecke 
/ Eing. Nr. 5, 1949” // “ZMH 827634”; “9” // “Type” // 
“Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // 
“pyroleuca 2” // “Sammlung / G Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827635”; “3” // “TYPE” // “Algeria mer. 
/ Djebel Aures / VII. 1913 /H. Stauder” // “Sammlung /G 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827636”. 

Original locality. Algeria: “in den algerischen Atlas 
und die nordliche Sahara” [in the Algerian Atlas and the 
northern Sahara]. 

Remarks. Stauder (1913) proposed this name as a 
form of 7! d. nouna (Lucas, 1848). Therefore, as stated 
by article 45.6.1 (CZN 1999) it is deemed to be an in- 
frasubspecific name (the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 7eracolus 
daira nouna Lucas, 1849 is junior subjective synonym of 
Colotis evagore nouna (Lucas, 1849). 


62. ab. evagorides Stauder, 1913 


Original combination. “Zeracolus daira nouna Luc. ab. 
evagorides m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290. 

Current combination. Colotis evagore nouna ab. 
evagorides (Stauder, 1913). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 2¢.¢ (ZMH 
827620-827621) (Fig. 62). “4” // “Algeria mer. / Djebel 
Aureés / VI. 1913/H. Stauder” // “Sammlung /G Warnecke 
/ Eing. Nr. 5,1949” // “COTYPE” // “ZMH 827620”; “3” 
// “evagorides / Stdr.” // “Algeria mer. / Djebel Aures / 
VII. 1912 / H. Stauder” // “TYPE” // “Sammlung / G 
Warnecke / Eing. Nr. 5,1949” // “ZMH 827621”. 

Original locality. Algeria: “in den algerischen Atlas 
und die nordliche Sahara” [in the Algerian Atlas and the 
northern Sahara]. 


Evolutionary Systematics 5 2021, 193-261 


Remarks. Stauder (1913) proposed this name as an 
aberation of 7’ d. nouna (Lucas, 1848). Therefore, as 
stated by article 45.6.1 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author expressly gave it 
infrasubspecific rank) and is hence unavailable. 7eracolus 
daira nouna Lucas, 1849 is junior subjective synonym of 
Colotis evagore nouna (Lucas, 1849). 


63. italorum Stauder, 1921 


Original combination. “Pieris brassicae L. f. n. (g. 
aest. mer. alticola) italorum” Stauder, 1921 Dt. Ent. Z. 
Iris 35: 26. 

Current combination. Pieris 
Stauder, 1921. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH 827617) (Fig. 63). 
“Calabria mer. / Aspromonte 900 m. / 29/6 Polsi [?] 1920 
/H. Stauder Legit.” // “Type” // “brassicae var. / italorum 
Staud.” // “ZMH 827617”. 

Type locality. Italy: “1.[talien] c.[entralis] m.[eridiona- 
lis], Aspromonte, Montalto, 1800 m.” 


brassicae italorum 


64. elbursina Bytinski-Salz & Brandt, 1937 


Original combination. “Pieris ergane, Hbn. ssp. e/bursina, 
ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: 1-2. 

Current combination. Pieris ergane elbursina By- 
tinski-Salz & Brandt, 1937. 

Current status. Valid subspecies. 

Type material. Syntypes 34'3' 19 (ZMH 827610- 
827613) (Fig. 64). “IRAN / Keredj (1500 m.) / 18/5 1936 
/ Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // 
“Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827610”; 
“TRAN / Keredj (1200 m.)/ 4.4. 1936 / Brandt” // “Cotypus 
/ P. ergane / ssp. pfeifferi / By.S & Br.” // “ssp. pfeifferi 
Byt.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 
827611”; “IRAN / Keredj (1500 m.) / 20/5 1936 / Brandt” 
//“Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “Coll By- 
tinski-Salz / Eing. Nr. 20, 1960” // ZMH 827612”; “IRAN / 
Keredj (1500 m.)/ 18/5 1936 / Brandt” // “Cotypus / ergane 
ssp. / pfeifferi / By.S+Br.” // “ssp. pfeifferi / By.S. & Br.” // 
“Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827613”. 

Type locality. Iran: Tehran, “Keredj” [Karaj]. 

Remarks. Label indicates ssp. pfeifferi, but apparent- 
ly this name does not exist. Collection data are identical 
to ssp. e/bursina, possibly a last-minute name change by 
Bytinski-Salz. 


65. r. exigua Verity, 1923 


Original combination. “Pieris ergane, H.-G. race exi- 
gua, mihi.” Verity, 1923 Ent. Rec. 35 Suppl. (18). 

Current combination. Pieris ergane r. exigua Veri- 
ty, 1923. 


217 


Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Cotype” 244 (ZMH 
835577—-835578) (Fig. 65). “exigua Vrty” // “Marche / 
Bolognola / 7 Ag 1937 / Querci” // [blank label] // “ZMH 
835577”; “exigua Vrty” // “Marche / Bolognola / 8 Ag 
1937 / Querci” // [blank label] // “ZMH 835578”. 

Original locality. Italy: “Upper Fargno Valley, at 
1400 m., in the Sibillini Mts.” 

Remarks. Verity (1923) proposed this name as a race 
of P. ergane (Geyer, 1828). According to Kudrna (1983), 
this name in unavailable because it is a seasonal form and 
also a nomen nudum. The year on the specimen labels 
(1937) indicate specimens were collected 14 years after 
description (1923) from the same location the original 
holotype material was collected. As a result, the black la- 
bel of the specimens treated as “types” is erroneous. The 
specimen collected from the same geographical location 
from which the type specimen was collected is called to- 
potype. According to ICZN (1999), a topotype has no for- 
mal standing and is not regulated by the Code. Therefore, 
these two specimens are erroneously labelled as types, 
should be annotated as “Not a Type” and are treated here 
as “Non-type” specimens. 


66. f. rossioides Stauder, 1921 


Original combination. “P. rapae L. f. n. rossioides” 
Stauder, 1921 Dt. Ent. Z. Iris 35: 27. 

Current combination. Pieris rapae f. rossioides 
Stauder, 1921. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827627) (Fig. 66). “Calabria mer. / Aspromonte 800 m. 
/ 4/7 [illegible] 1920 / H. Stauder legit.” // “Type” 
// “Rossioides Stdr.” // “rossioides / Stdr. / Type” // 
“ZMH 827627”. 

Original locality. Italy: “Polsibeken, Aspromonte 
7-900 m.” 

Remarks. Stauder (1921) proposed this name as a 
form of P. rapae (Linnaeus, 1758). According to article 
45.6.1 (ICZN 1999), it is infrasubspecific if the content 
of the work (please see below) unambiguously reveals 
that the name was proposed for an infrasubspecific en- 
tity. In the same paper, Stauder (1921) described another 
“form” of P. rapae as “zelleri” and clearly stated that it 
also occurs at the same “type locality” as rossioides (1.e., 
Aspromonte). As by definition two geographical races (or 
forms) cannot occur sympatrically; therefore, the content 
of the work reveals that such forms were not meant to be 
subspecific. However, Stauder added further complexity; 
there is a difference between how “zelleri’” and “rossioi- 
des” have been described. In fact, the former was as ‘g. 
aest. mer. alt.’, 1.e., generatio aestivalis meridionalis alti- 
tudinalis [Southern altitudinal summer generation], while 
the latter just as ‘f. n.’, 1.e., “forma nova” [new form]. 


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218 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Pn te : fas 
61a Gaae | Sue 


ZMH 827618| 


7 


hee ee 


‘Coli ee 
Eimg_ Nr. 20, 1960 


ZMH_ 835577 


1cm 


Plate 10. 59. Colotis evagore nouna f. vernalis alticola auresiaca (Stauder, 1913); 60. Colotis evagore nouna f. biformata (Stauder, 
1913); 61. Colotis evagore nouna f. aestiva pyroleuca (Stauder, 1913); 62. Colotis evagore nouna ab. evagorides (Stauder, 1913); 
63. Pieris brassicae italorum Stauder, 1921; 64. Pieris ergane elbursina Bytinski-Salz & Brandt, 1937; 65. Pieris ergane r. exigua 
Verity, 1923. a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


Thus, it could theoretically be argued that rossioides was 
meant as a geographical race, while zelleri would be a 
seasonal variation of that. However, this speculation does 
not seem to be correct, because it makes no sense that 
someone described a seasonal form of a subspecies be- 
fore describing the subspecies itself. As a result, looking 
at the paper as a whole and not to the isolated description 
of rossioides alone, this name has to be considered as in- 
frasubspecific (Alberto Zilli pers. comm. ). 


Unpublished, unavailable names 


67. ab. gottingensis Meinhard 


Original combination. Pieris napi v. napaeae ab. got- 
tingensis Meinhard. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Type” 14 (ZMH 
835579) (Fig. 67). “v. gottingensis / Mht” // “8/5 1909 / 
Gottingen / Hainholz” // “ZMH 835579”. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 


68. ab. nigromarginata Meinhard 


Original combination. Pieris brassicae ab. nigromar- 
ginata Meinhard. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Type” 14 (ZMH 
835576) (Fig. 68). “13/8 1916 / Gottingen / Hainberg” // 
“ZMH 835576”. 

Remarks. This taxon cannot be found in the literature. 
As Stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and 1s hence unavailable. 


Family Lycaenidae Leach, 1815 
Subfamily Aphnaeinae Distant, 1884 


69. dueldueli Pfeiffer, 1932 


Original combination. “Cigaritis acamas Klug. ssp. n. 
dildiili m.” Pfeiffer, in Osthelder and Pfeiffer 1932, Mitt. 
Munch. Ent. Ges. 22: 38. 

Current combination. Cigaritis acamas dueldueli 
Pfeiffer, 1932. 

Current status. Valid subspecies. 

Type material. Syntypes 3¢¢ (ZMH 827553- 
827555) (Fig. 69). “Syria s. / amanus / Duldtl Dagh 
/ VII.30” // “Cotypus / A. epargyros / ssp. diildiili / 
Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” 
// ZMH 827553”; “Syria sept. / amanus sept. / Duldiul 
Dagh / VII.30” // “Cotypus / A. epargyros / ssp. diildiili 


219 


/ Pfeiffer” // “Coll. Bytinski Salz / Eing. Nr. 20, 1960” 
/| “ZMH 827554”; “Syria sept. / amanus sept. / Duldtl 
Dagh / VH.30” // “Cotypus / A. epargyros / ssp. diildiili 
/ Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // 
“ZLMEP8273599" - 

Type locality. Turkey: “Marasch in tirkisch Nordsy- 
rien; Im DD [Dildtl Dagh].” 


Subfamily Lycaeninae Leach, 1815 
Tribe Lycaenini Leach, 1815 


70. f. emarginata Warnecke, 1942 


Original combination. “Chrysophanus  alciphron 
Rott., J, n. f emarginata.” Warnecke, 1942 Dt. Ent. Z. 
Iris 56: 103. 

Current combination. Lycaena alciphron f. emargi- 
nata (Warnecke, 1942). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827527) (Fig. 70). “Schlesw. — Holst. / Kr. / Bockhorn / 
W. Wolf leg. / 14.7.40” // “Katalog Nr. / L 2322” //“¢ nf. 
emarginata / Warn. Type / Iris 1942, 103” // “Chrysopha- 
nus | alciphron / emarginata Warn. / Type / L.31” // “Sig. 
G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827527”. 

Originallocality. Germany: “Segeberg’”; Schleswig-Hol- 
stein publication says Segeberg; the label of the specimen 
treated as type by Warnecke states Lauenburg crossed out 
and replaced by Bockhorn. 

Remarks. Warnecke (1942) proposed this name as a 
form of C. alciphron (Rottemburg, 1775). According to 
article 45.6.1 (ICZN 1999), it is infrasubspecific if the 
content of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity, and is hence 
unavailable. Already the title of Warnecke (1942) paper 
reveals that this form was described as an aberration and 
not as a subspecies. 


71. ab. alba Schoenfeld, 1924 


Original combination. “Chrysophanus virgaureae L.-9, 
alba m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40. 

Current combination. Lycaena virgaureae ab. alba 
(Schoenfeld, 1924). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 12 (ZMH 
827586) (Fig. 71). “Bayern / Mitterteich / 24. VHI.1923 / 
Schonfeld” // [blank label] // “Type / Chr. virgaureae L. / 
ab. alba Schonfeld /TI. E. Z. Guben 18. Jahrg / 1924. Nr 6. 
P. 40 / [illegible]” // “ZMH 827586”. 

Original locality. Germany: Bavaria. 

Remarks. Schoenfeld (1924) proposed this name as 
an aberration of C. virgaureae (Linnaeus, 1758), there- 
fore, as stated by article 45.6.2 (ICZN 1999) it is deemed 
to be an infrasubspecific name (the author used “aberra- 
tion’, “ab.”) and is hence unavailable. 


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220 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


72. f. multipunctata Warnecke, 1942 


Original combination. “Chrysophanus virgaureae L., 3, 
n.f. multipunctata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103. 

Current combination. Lycaena virgaureae f. multi- 
punctata (Warnecke, 1942). 

Current status. Infrasubspecific and hence unavailable. 

Original material. Labelled as “Type” 14 (ZMH 
827592) (Fig. 72). “n.f. multipunctata / Warn. Type / Iris 
1942, 103” //“Chrysoph. virgaureae / multipunctata / Type 
Warn. / L.30” // “Holstein” // “Katalog Nr. / L. 2196” // 
“Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827592”. 

Original locality. Germany: Holstein. 

Remarks. Warnecke (1942) proposed this name as a 
form of C. virgaureae (Linnaeus, 1758). According to 
article 45.6.1 (ICZN 1999), it is infrasubspecific if the 
content of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity, and is hence 
unavailable. The title and the text of Warnecke (1942) pa- 
per unambiguously reveals that this form was described 
as an aberration and not as a subspecies. It was only found 
in a single male. 


73. f. quercii Turati, 1923 


Original combination. “Chrysophanus virgaureae emil- 
ianus f.n. e. quercii f. n.” Turati, 1923 Atti. Soc. Ital. Sci. 
Nat. 62: 42. 

Current combination. Lycaena virgaureae emil- 
ianus f. quercii (Turati, 1923). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 646 (ZMH 
827580-827585) (Fig. 73). “Marche / Bolognola / 
15.VI.1937 / Querci” // [blank label] // “quercii Trti.” 
/1 “ZMH 827580”; “Marche / Bolognola / 15. VII.1937 / 
Querci” // [blank label] //“quercii Trti.” // “ZMH 827581”; 
“Marche / Bolognola / 3 Ag 1937 / Querci” // [blank la- 
bel] // “quercii Trti.” // “ZMH 827582”; “Marche / Bo- 
lognola / 27. VII.1937 / Querci” // [blank label] // “quer- 
cil Trti.” // “ZMH 827583”; “Marche / Bolognola / 5 Ag 
1937 / Querci” // [blank label] // “quercui Trti.” // “ZMH 
827584”; “Marche / Bolognola / 4 Ag 1937 / Querci” // 
[blank label] // “quercit Trti.” // “ZMH 827585”. 

Original locality. Italy: “Monti Sibillini’. 

Remarks. Turati (1923) proposed this name as a 
form of C. v. emilianus Turati, 1923. Therefore, as stat- 
ed by article 45.6.1 (ICZN 1999) it is deemed to be an 
infrasubspecific name (the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 


Subfamily Polyommatinae Swainson, 1827 
Tribe Polyommatini Swainson, 1827 


74. andarabi Forster, 1938 


Original combination. “Lycaena pheretiades andarabi 
ssp. nov.” Forster, 1937 Mitt. Minch. Ent. Ges. 27: 61. 


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Current combination. Agriades pheretiades andarabi 
(Forster, 1938). 

Current status. Valid subspecies. 

Type material. Paratypes 4¢6 (ZMH 827575- 
827578) (Fig. 74). “West-Hindukusch / Andarab / Gerdoll- 
zone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” 
// “Para-Type / Lyc. Pheretiades / andarabi Forst. / W. 
Forster Munchen” // “Coll. Bytinski Salz / Eing. Nr 20, 
1960” // “ZMH 827575”; “Nord-Ost-Hindukusch / Nuk- 
san-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m 
Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. 
Pheretiades / andarabi Forest. / W. Forster Munchen” 
// “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 
827576”; “West-Hindukusch / Andarab / Gerollzone / 
4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Pa- 
ra-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster 
Munchen” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // 
“ZMH 827577”; “West-Hindukusch / Andarab / Geroll- 
zone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” 
// “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. 
Forster Munchen” // “Coll. Bytinski-Salz / Eing. Nr 20, 
1960” // “ZMH 827578”. 

Type locality. Afghanistan: “West-Hindukusch, Anda- 
rab, 4000-4500 m.” 


75. f. altera Ziillich, 1929 


Original combination. “Lycaena anteros Frr. f. altera” 
Zullich, 1929 Z. Ost. Ent. Ver. 14: 52. 

Current combination. Arvicia anteros f. altera (Ziil- 
lich, 1929). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type’1¢ (ZMH 
827526) (Fig. 75). “BULGARIA OR. / GIOKTEPE 250 
M/ MITTE AUG 1928 / COLL. ZUELLICH” // “ExTyp- 
icalibus / Specimenibus” // “VIHI 1928 / Gidktepe, / Bul- 
garien” // “ZMH 827526”. 

Original locality. Bulgaria: “Rilogebiet, Sidwestbul- 
garien, 1200 m, und Gioktepe, Stidostbulgarien, 200 m.” 

Remarks. Zillich (1929) proposed this name as a 
form of L. anteros Freyer, 1839. According to article 
45.6.1 (ICZN 1999), it is infrasubspecific if the content 
of the work unambiguously reveals that the name was 
proposed for an infrasubspecific entity, and 1s hence un- 
available. The text of Zullich (1929: 52) unambiguously 
reveals that this form was described explicitely for the 
summer generation. 


76. r. modicior Verity, 1938 


Original combination. “Aricia anteros, Frr. modicior, 
nom. nov.” Verity, 1938 Ent. Rec. 50(8): Suppl. (5). 
Current combination. Aricia anteros r. modicior 
Verity, 1938. 
Current status. Infrasubspecific and hence unavail- 
able name. 


Evolutionary Systematics 5 2021, 193-261 221 


ZMH 835579 


ZMH ‘8276271 


ociceesy 
[ZMH 827527| 


ZMH -827586 
ZMH -827592| 


i1cm 


Plate 11. 66. Pieris rapae f. rossioides Stauder, 1921; 67. Pieris napi v. napaeae ab. gottingensis Meinhard; 68. Pieris brassicae ab. 
nigromarginata Meinhard; 69. Cigaritis acamas dueldueli Pfeiffer, 1932; 70. Lycaena alciphron f. emarginata (Warnecke, 1942); 


71. Lycaena virgaureae ab. alba (Schoenfeld, 1924); 72. Lycaena virgaureae f. multipunctata (Warnecke, 1942). a. Dorsal view, 
b. ventral view. 


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222 


Original material. Labelled as “Type” 233’, (ZMH 
827535—-827536) (Fig. 76). “modicior Vrty” // “Macedo- 
nia / Olympus, 4500’ / July 2, 1935 / Romev” // [blank la- 
bel] // “ZMH8274535”; “modicior Vrty” // “anteros Frr.” 
// “Macedonia / Olympus, 4500’ / July14, 1935 / Rome1” 
// [blank label] // “ZMH8274536”. 

Original locality. Greece: “At the highest altitudes of 
Mt. Olympus, such as at Prionia, 1500 m”. 

Remarks. According to Kudrna (1983: 32), this name 
is unavailable and denotes a race and seasonal form and 
is hence unavailable. 


77. uliginosa Dannehl, 1921 


Original combination. “Lycaena argus Schiff. var. uligi- 
nosa” Dannehl, 1921 Mitt. Minch. Ent. Ges. 11: 39. 

Current combination. Plebejus idas uliginosa (Dan- 
nehl, 1921). 

Current status. Junior subjective synonym of Plebe- 
Jus argus argus (Linnaeus, 1758). 

Type material. Syntypes 443, (ZMH 827541- 
827542, ZMH 835595-835596) (Fig. 77). “18.7” // 
“Ob. Bayern / Beuerberg / c. Dannehl 20” // “Argus v. 
n. / uliginosa” // “Katalog Nr. / L 2562” // “Cotype” // 
“L.34/Lyc. argus -/v. uliginosa Dannehl / 3 Cotype” // 
Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827541”: 
“18.7” // “Ob. Bayern / Beuerberg / c. Dannehl 20” // 
“vy. uliginosa / Dannehl” // “Cotype” // “Katalog Nr. / 
L 2561” // “L.35 / Lye. argus - / v. uliginosa Dannehl 
/ 2 Cotype” // Sig. G. Warnecke / Eing. Nr. 5, 1949” 
// “ZMH 827542”; “27.6” // Ob. Bayern / Beuerberg 
/ F. Dannehl” // “Cotype” // “27.6. 1921 / Beuerberg 
/ Oberbayern / v. Dannehl” // “ZMH 835595”; “13.7” 
// Ob. Bayern / Beuerberg / F. Dannehl” // “Cotype” // 
“13.7. 1921 / Beuerberg / Oberbayern / v. Dannehl” // 
“LMIEU8 35596". 

Type locality. Germany: Bavaria, Beuerberg. 

Remarks. Dannehl (1921) proposed this name as a 
“var.” of L. argus ({[Denis & Schiffermitiller], 1775). L. ar- 
gus 1S a junior subjective synonym of Plebejus idas Lin- 
naeus, [1760]. According to article 45.6.4 ICZN 1999), 
it is subspecific if first published before 1961 and author 
expressly used the term “var.”. 


78. balcanica Ziillich, 1929 


Original combination. “Lycaena argyrognomon Bgstr. f. 
balcanica” Zillich, 1929 Z. Ost. Ent. Ver. 14: 52. 

Current combination. Plebejus argyrognomon bal- 
canica (Ziillich, 1929). 

Current status. Preoccupied by Lycaena semiargus 
balcanica Tutt, 1909; replacement name Lycaeides ar- 
gyrognomon baldur (Hemming, 1934) is the current val- 
id name. 

Type material. Holotype 14 (ZMH 827544) 
(Fig. 78) paratype 146 (ZMH 827543). “BULGARIA 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


OCC. // MON. RILSKI 1200 M / 20. — 31 VII. 1928 
// COLL. ZUELLICH” // “Ex Typicalibus / Specimini- 
bus” // “Holotype” // “Coll. Bytinski-Salz / Eing Nr 20, 
1960” // ZMH 827544”; “BULGARIA OCC. // MON. 
RILSKI 1400 M / 20 VI. — 6 VII. 1928 // COLL. ZU- 
ELLICH” // “Ex Typicalibus / Speciminibus” // “Para- 
typoid” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // 
ZMH 827543”. 

Type locality. Bulgaria: “MON. RILSKI” [Monas- 
tery Rilski]. 

Remarks. Zillich (1929) proposed this name as a 
form of L. argyrognomon (Bergstrasser, 1779). Accord- 
ing to article 45.6.4 (ICZN 1999), it is subspecific if first 
published before 1961 and author expressly used the 
term “form”. 


79. mayencis Eitschberger & Steiniger, 1975 


Original combination. “Eumedonia eumedon mayencis n. 
subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (1): 106. 

Current combination. Eumedonia eumedon may- 
encis Eitschberger & Steiniger, 1975. 

Current status. Valid subspecies. 

Type material. Paratypes 546 (ZMH 827548-— 
827552) (Fig. 79). “Gonsenheim / 5. Juni / 04 / AN- 
DREAS / Gonsenheim” // “PARATYPUS 0 / Eumedonia 
eu- / medon mayencis | EITTSCHBERGER & / STEINI- 
GER, 1975” //“Coll. Bytinski Salz / Eing. Nr 20, 1960” // 
“ZMH 827548”; “Gonsenheim / 5. Juni / 04 / ANDREAS 
/ Gonsenheim” // “PARATYPUS o / Eumedonia eu- / 
medon mayencis | ETTSCHBERGER & / STEINIGER, 
1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” //“ZMH 
827549”; “Mainz / Mombach / 10.6.1922” // “Eumedon” 
// “PARATYPUS o / Eumedonia eu- / medon mayencis 
/ EITSCHBERGER & / STEINIGER, 1975” // “Coll. 
Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827550”; 
“Gonsenheim / 8. Juni / 04 /ANDREAS / Gonsenheim” // 
“PARATYPUS 0 / Eumedonia eu- / medon mayencis / EI- 
TSCHBERGER & / STEINIGER, 1975” // “Coll. Bytins- 
ki Salz / Eing. Nr 20, 1960” // “ZMH 827551”; “Mainz / 
Mombach / 10.6.1922” // “Eumedon” // “PARATYPUS o 
/ Eumedonia eu- | medon mayencis | ETYTSCHBERGER 
& / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 
20, 1960” // “ZMH 827552”. 

Type locality. Germany: “Deutschland, Rhein- 
land-Pfalz, Mainz-Mombach, Mainzer Sand”. 


80. moenus Eitschberger & Steiniger, 1975 


Original combination. “Eumedonia eumedon moenus n. 
subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (2): 99. 

Current combination. Eumedonia eumedon moenus 
Eitschberger & Steiniger, 1975. 

Current status. Valid subspecies. 

Type material. Paratypes 2¢¢ (ZMH 827546-— 
827547) (Fig. 80). “Tiertalsberg be1/ Retzbach am Main 


Evolutionary Systematics 5 2021, 193-261 


/ 4.6.1950” // “Katalog Nr. / L 3370” // “PARATYPUS 
o / Eumedonia eume- / don moenus | EITSCHBERG- 
ER & / STEINIGER, 1975” // “Bot.-Zool. Excursion 
/ 1950/ Eing Nr. 14—1950” // “ZMH 827546”; “Karl- 
stadt a. Main / Karlsburg / 4.6.1950” // “Katalog Nr. / 
L 3369” // “PARATYPUS 0 / Eumedonia eume- / don 
moenus | EITSCHBERGER & / STEINIGER, 1975” // 
“Bot.-Zool. Excursion / 1950/ Eing Nr. 14—1950” // 
“ZMH 827547”. 

Type locality. Germany: “Deutschland, Nordbayern, 
mittleres Maintal und Nord-Baden, mittleres Taubertal.” 


81. persa Bytinski-Salz & Brandt, 1937 


Original combination. “Chilades trochylus, Frr. ssp. 
persa, ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 
49: Suppl. (2). 

Current combination. Freyeria trochylus persa (By- 
tinski-Salz & Brandt, 1937). 

Current status. Valid subspecies. 

Type material. Syntypes 4¢¢ (ZMH 827587-— 
827590) (Fig. 81). “IRAN / Keredj (1600 m.) / 3/6 1936 
/ Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S 
+ Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // 
“ZMH 827587”; “IRAN / Keredj (1600 m.) / 9/8 1936 
/ Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S 
+ Brandt” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // 
“ZMH 827588”; “IRAN / Keredj (1600 m.) / 12/8 1936 
/ Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S 
+ Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // 
“ZMH 827589”; “IRAN / Keredj (1600 m.) / 3/6 1936 
/ Brandt” // “Cotypus / Ch. trochylus / ssp. persa / ByS 
+ Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // 
“ZMH 827590”. 

Type locality. Iran: Tehran, “Keredj” [Karaj]. 


82. ab. pauper Bytinski-Salz & Brandt, 1937 


Original combination. “Chilades trochylus, Frr. ssp. 
persa, ssp. nov. ab. pauper, ab. nov.” Bytinski-Salz & 
Brandt, 1937 Ent. Rec. 49: Suppl. (2). 

Current combination. Freyeria trochylus persa ab. 
pauper (Bytinski-Salz & Brandt, 1937). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Holotype” 9 (ZMH 
827591) (Fig. 82). “IRAN / Keredj (1600 m.) / 25/5 1936 
/ Brandt” // “ab. pauper / ByS+Br. / Holotype 9 / ex. coll. 
/ Dr. H. Bytinski-Salz” // “Coll. Bytinski-Salz / Eing. Nr. 
20, 1960” // “ZMH 827591”. 

Original locality. Iran: Tehran, “Keredj” [Karaj]. 

Remarks. Bytinski-Salz and Brandt (1937) proposed 
this name as an aberration of C. trochylus (Freyer, 1838). 
Therefore, as stated by article 45.6.2 ICZN 1999) it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”) and is hence unavailable. 


223 


83. parvandereggi Verity, 1938 


Original combination. “Glaucopsyche cyllarus, Rott, 
race parvandereggi, nom. nov.” Verity, 1938 Ent. Rec. 
50: Suppl. (4). 

Current combination. Glaucopsyche alexis parvan- 
dereggi Verity, 1938. 

Current status. Valid subspecies. 

Type material. Syntypes 246 (ZMH 833470- 
833471) (Fig. 83). “parvandereggi Vrty” // “Macedonia 
/ Olympus, 2900’ / V. 25, 1936 / Romei” // [blank label] 
// “ZMH 833470”; “parvandereggi Vrty” // “Macedonia / 
Olympus, 2900’ / V. 26, 1936 / Romei” // [blank label] // 
“ZMH 833471”. 

Type locality. Greece: Salonika [Macedonia, Olympus]. 

Remarks. Verity (1938) proposed this name as a race 
of G. cyllarus (Rottemburg, 1775). As it was given in 
trinomy and the content of the description clearly indi- 
cates that a geographical race was meant, this name 1s 
fully available as subspecific (article 45.6 ICZN 1999). 
(Alberto Zilli pers. comm. ). 


84. sibyllina Verity, 1916 


Original combination. “Lycaena coridon” Poda, “razza 
sibyllina” Verity, 1916 Boll. Soc. Ent. Ital. 46: 133. 

Current combination. Lysandra coridon sibyllina 
(Verity, 1916). 

Current status. Junior subjective synonym of Lysan- 
dra coridon apennina (Zeller, 1847). 

Original material. Labelled as “Type” 404 (ZMH 
833493-833496) (Fig. 84). “sibyllina Vrty” // “Marche / 
Bolognola / 15 Ag 1937 / Querci” // [blank label] // “ZMH 
833493”; “sibyllina Vrty” // “Marche / Bolognola/9 Ag 1937 
/ Querci” // [blank label] // “ZMH 833494”; “sibyllina Vrty” 
//““Marche / Bolognola / 9 Ag 1937 / Querci” // [blank label] 
/!“ZMH 833495”; “sibyllina Vrty” // “Marche / Bolognola / 
8 Ag 1937 / Querci” // [blank label] // “ZMH 833496”. 

Original locality. Italy: “Monti Sibillini (Bolognola, 
1200 m)” 

Remarks. Verity (1938) proposed this name as a “raz- 
za’ (=race) of L. coridon (Poda, 1761). As it was given in 
trinomy and the content of the description clearly indicates 
that a geographical race was meant, this name is avail- 
able as subspecific (article 45.6 ICZN 1999). However, it 
was later synonimized with L. coridon apennina (Zeller, 
1847). The year on the specimen labels (1937) indicate 
specimens were collected 21 years after description (1916) 
from the same location the original holotype material was 
collected. As a result, the black label of the specimens 
treated as “types” is erroneous. The specimen collected 
from the same geographical location from which the type 
specimen was collected is called topotype. According to 
ICZN (1999), a topotype has no formal standing and is not 
regulated by the Code. Therefore, these four specimens are 
erroneously labelled as types, should be annotated as “Not 
a Type” and are treated here as “Non-type” specimens. 


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224 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Coll. Bytinghi Salz 


fom 


ornate Ka widsa 
e.Daunchl | 


esas 


y Te 
WS Mt 
k 
¢ 
/ Eo 
at. 
oe 
cP 


77b- 


SS ndaeaniane mae 


- Coll. Bytingsi Saiz 
| Eng. Nr. 29, _ 1969 


» lar ara 


79a 


ZMH 827550 


we eeeeeee 


yy 


' ytinski-Salz 


: ew 
ing. We, 70. 1960 fine Nr. 20, 1969 


| ene. enn 2MH - 827589 ZMH 827591 


[zm 827546| 


ZMH _ 833493 


ZMH 833470 


1cm 


Plate 12. 73. Lycaena virgaureae emilianus f. quercii (Turati, 1923); 74. Agriades pheretiades andarabi (Forster, 1938); 75. Aricia 
anteros f. altera (Zullich, 1929); 76. Aricia anteros r. modicior Verity, 1938; 77. Plebejus idas uliginosa (Dannehl, 1921); 78. Plebejus 
argyrognomon balcanica (Zillich, 1929); 79. Eumedonia eumedon mayencis Eitschberger & Steiniger, 1975; 80. Eumedonia eum- 
edon moenus Eitschberger & Steiniger, 1975; 81. Freyeria trochylus persa (Bytinski-Salz & Brandt, 1937); 82. Freyeria trochylus 
persa ab. pauper (Bytinski-Salz & Brandt, 1937); 83. Glaucopsyche alexis parvandereggi Verity, 1938; 84. Lysandra coridon sibyl- 
lina (Verity, 1916). a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


85. f. lunulata Warnecke, 1942 


Original combination. “Lycaena alcon F., 9, n. f. 
lunulata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103. 

Current combination. Phengaris alcon f. lunulata 
(Warnecke, 1942). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 12 (ZMH 
827528) (Fig. 85). “Bahrenfeld / 28.7.08” // “Sig. S. War- 
necke / Eing. Nr. 5 1949” // “9 nf. unuata / Warn. Type / 
Iris 1942, 103” // “Katalog Nr. L 32 67” // “ZMH 827528”. 

Original locality. Germany: Eidelstedt, Hamburg. 

Remarks. Warnecke (1942) proposed this name as 
a form of “ZL. alcon F.” (sic, recte: [Denis & Schiffer- 
miller], 1775). According to article 45.6.1 (ICZN 1999), 
it is infrasubspecific if the content of the work unambig- 
uously reveals that the name was proposed for an infra- 
subspecific entity, and is hence unavailable. The title of 
the paper (Warnecke, 1942) indicates that this form con- 
stitutes an aberration and is therefore infrasubspecific and 
unavailable. According to the description this form was 
only found in a single female. 


86. f. selzeri Warnecke, 1924 


Original combination. “Lycaena arionides Staudinger f. 
selzeri” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149. 

Current combination. Phengaris arionides f. selzeri 
(Warnecke, 1924). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827545) (Fig. 86). “Ussuri” // “F. selzeri Warn. / 3 Type” 
// “nf. selzeri Warn. / I. Iris 1924, 38. J. / p. 149, Taf. I 
bis. 5.” // “Type” // “Lyc. arionides - / n.f. selzeri Warn. 
/ L.33” // “Katalog Nr. / L 3336” // “Sig. G. Warnecke / 
Eing. Nr. 5, 1949” // “ZMH 827545”. 

Original locality. Russia: “Ostsibirien, Amur-Gebiet, 
Ussuri”. 

Remarks. Warnecke (1942) proposed this name as a 
form of L. arionides Staudinger, 1887. According to arti- 
cle 45.6.1 (ICZN 1999), it is infrasubspecific if the content 
of the work unambiguously reveals that the name was pro- 
posed for an infrasubspecific entity, and is hence unavail- 
able. According to the description this form is an individ- 
ual aberration or a rare second generation and is therefore 
infrasubspecific. Accordingly, on plate II this form is fig- 
ured as “Lycaena arionides nov. ab. selzeri Warn.”. 


87. peninsulae Verity, 1923 


Original combination. “Lycaena euphemus, Hub. (a) race 
peninsulae, mihi” Verity, 1923 Ent. Rec. 35 Suppl. (7). 

Current combination. Phengaris euphemus penin- 
sulae (Verity, 1923). 


225 


Current status. Valid subspecies. 

Original material. Labelled as “Type” 14 (ZMH 
833468) (Fig. 87). “peninsulae Vrty” // “Marche / Bolog- 
nola / 10.VI.1937 / Querci / 1300 m” // [blank label] // 
“ZMH 833468”. 

Original locality. Italy: “Sibillini Mts. in the Marche, 
below Bolognola, at 1000 m”. 

Remarks. Verity (1923) proposed this name as a race 
of L. euphemus Hubner, 1799-1800. According to Kudrna 
(1983: 35), this race represents an available subspecific 
name. However, the year on the specimen labels (1937) 
indicate specimens were collected 14 years after descrip- 
tion (1923) from the same location (..e., Italy: Bolognola) 
the original type material was collected. As a result, the 
black label of the specimens treated as “type” by Veri- 
ty is erroneous. The specimen collected from the same 
geographical location from which the type specimen was 
collected is called topotype. According to ICZN (1999), 
a topotype has no formal standing and is not regulated 
by the Code. Therefore, this specimen is erroneously la- 
belled as type, should be annotated as “Not a Type” and 
are treated here as “Non-type” specimen. 


88. ab. depuncta Hirschke, 1903 


Original combination. “Lycaena Argiades ab. (et var.) 
depuncta” Hirschke, 1903 Verh. Ges. Wien. 53: 270. 

Current combination. Cupido argiades ab. depunc- 
ta (Hirschke, 1903). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827531) (Fig. 88). “Hirschke.Wien / Herkulsbad / V.1900 
Type” // “Verh. zool. / bot. ges. / 1903 Hirs.” // “TYPE” 
/1 “ZMH 827531”. 

Original locality. Romania: “Herkulesbad”. 

Remarks. Hirschke (1903) proposed this name as an 
aberration of L. argiades (Pallas, 1771). Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


89. ausonia Verity, 1913 


Original combination. “Lycaena damon ausonia nom. 
nov.” Verity, 1913 Boll. Soc. Ent. Ital. 45: 232, pl. 1, f. 
44,47. 

Current combination. Polyommatus (Agrodiaetus) 
damon ausonia (Verity, 1913). 

Current status. Junior subjective synonym of Poly- 
ommatus (Agrodiaetus) damon damon ({Denis & Schif- 
fermuller], 1775). 

Original material. Labelled as “Type” 63'¢ (ZMH 
833479-833484) (Fig. 89). “ausonia Vrty.” // “Marche 
/ Bolognola / 15.VII.1937 / Querci” // [blank label] // 
“ZMH 833479”; “ausonia Vrty.” // “Marche / Bolognola / 


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226 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


15.VH.1937 / Querci” // [blank label] // “ZMH 833480”; 
“ausonia Vrty.” // “Marche / Bolognola / 12.VII.1937 
/ Querci” // [blank label] // “ZMH 833481”; “auso- 
nia Vrty.” // “Marche / Bolognola / 5 Ag 1937 / Quer- 
ci” // [blank label] // “ZMH 833482”; “ausonia Vrty.” // 
“Marche / Bolognola/3 Ag 1937 / Querci” // [blank label] 
//“ZMH 833483”; “ausonia Vrty.” // “Marche / Bolognola 
/ 12. VII.1937 / Querc1” // [blank label] // “ZMH 833484”. 

Original locality. Italy: “a grandi altitudini nei soliti 
monti del Piceno” [At high elevations at Mount del Piceno]. 

Remarks. The year on the specimen labels (1937) in- 
dicate specimens were collected 24 years after descrip- 
tion (1913) from the same location the original holotype 
material was collected. As a result, the black label of the 
Specimens treated as “types” is erroneous. The speci- 
men collected from the same geographical location from 
which the type specimen was collected is called topo- 
type. According to ICZN (1999), a topotype has no for- 
mal standing and is not regulated by the Code. Therefore, 
these six specimens are erroneously labelled as types, 
should be annotated as “Not a Type” and are treated here 
as “Non-type” specimens. 


90. pfeifferi Brandt, 1938 


Original combination. “Lycaena Pfeifferi sp. n.” Brandt, 
1938 Ent. Rdsch. 55: 671, pl. 4, f. 1-3. 

Current combination. Polyommatus (Agrodiaetus) 
Dfeifferi (Brandt, 1938). 

Current status. Valid species. 

Type material. Paratype 14 (ZMH 827579) (Fig. 
90). “IRAN. FARS, StrafBe / Ardekan — Talochosroe / 
Comé, m, 1937 / coll. Brandt” //“L.37 / Paratypoid / Lyc. 
Dfeifferi Brand” // “Katalog Nr. / L. 3190” // “Sammlung 
R. Jahnig / Eing. Nr.29, 1949” // “ZMH 827579”. 

Type locality. Iran: Fars “vom Barm I Firus (3750 m)” 
[from Barm-e Firuz]. 


91. alcestis Zerny, 1932 


Original combination. “Lycaena (Hirsutina) ripperti 
alcestis Zerny nov. subsp.” Zerny, 1932 Dt. Ent. Z. Iris 
46: 186. 

Current combination. Polyommatus (Agrodiaetus) 
alcestis (Zerny, 1932). 

Current status. Valid species. 

Type material. Syntypes 2¢¢ (ZMH 827529- 
827530) (Fig. 91). “Syria / Libanon s. / Bscharre / VIII- 
IX 31 / 1300 m / Einh.Slir.leg” // “Cotypus / ripperti / 
ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 20, 
1960” // “ZMH 827529”; “Syria / Libanon s. / Bscharre / 
VIU-IX 31 / 1300 m/ Einh.Slrleg” // “Cotypus / ripperti 
/ ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 
20, 1960” // “ZMH 827530”. 

Type locality. “Lebanon, Becharré, 1400 m; Zedern- 
wald oberhalb Becharré um 1900 m”. 


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92. ardschira Brandt, 1938 


Original combination. “Lycaena ardschira sp. n.” 
Brandt, 1938 Ent. Rdsch. 55: 672. 

Current combination. Polyommatus (Agrodiaetus) 
ardschira (Brandt, 1938). 

Current status. Valid species. 

Type material. Paratype 1, (ZMH 827532) (Fig. 92). 
“L.36 / Paratypoid / Lyc. ardschira / Brand” // “Paratyp- 
oid” // “IRAN. FARS, StraBe / Ardekan — Talochosroe / 
Comé, m, 1937 /coll. Brandt” // “196” // “Sammlung / R. 
Jahnig / Eing. Nr. 29,1949” // “Katalog Nr. / L. 3365” // 
“ZMH 827532”. 

Type locality. Iran: Fars, “Comée, 2600 m” [Komehr]. 


93. f. squalida Verity, 1920 


Original combination. “Polyommatus meleager, Esp., 
race macra, mihi, 2 form squalida, mihi” Verity, 1920 
Ent. Rec. 32:145. 

Current combination. Polyommatus daphnis macra 
f. squalida Verity, 1920. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 24: (ZMH 
833485—833486) (Fig. 93). “squalida Vrty” // “Marche 
/ Bolognola / 15.VII.1937 / Querci” // [blank label] // 
“ZMH 833485”; “squalida Vrty” // “Marche / Bolognola 
/3 Ag 1937 / Querci” // [blank label] // “ZMH 833486”. 

Original locality. Italy: “Mt. Sibillina, 1200-1300 m”. 

Remarks. Verity (1920) proposed this name as a form 
of P. meleager macra Verity, 1920. According to Kudrna 
(1983: 39), this name is unavailable and denotes a race 
and is infrasubspecific and hence unavailable. Papilio 
meleager Esper, 1779 is a junior subjective synonym of 
Papilio daphnis {Denis & Schiffermiuller], 1775. The year 
on the specimen labels (1937) indicate specimens were 
collected 17 years after description (1920) from the same 
location the original holotype material was collected. As a 
result, the black label of the specimens treated as “types” 
is erroneous. The specimen collected from the same geo- 
graphical location from which the type specimen was 
collected is called topotype. According to ICZN (1999), 
a topotype has no formal standing and is not regulated by 
the Code. Therefore, these two specimens are erroneous- 
ly labelled as types, should be annotated as “Not a Type” 
and are treated here as “Non-type” specimens. 


94. r. correpta Verity, 1920 


Original combination. “Agriades hylas, Esp., race cor- 
repta, mihi” Verity, 1920 Ent. Rec. 32: 144. 

Current combination. Polyommatus dorylas r. cor- 
repta (Verity, 1920). 

Current status. Infrasubspecific and hence unavail- 
able name. 


Evolutionary Systematics 5 2021, 193-261 


Original material. Labelled as “Type” 40:40 (ZMH 
833489-833492) (Fig. 94). “correpta Vrty” // “Marche 
/ Bolognola / 15.VII.1937 / Querci / 1500 m” // [blank 
label] // “ZMH 833489”; “correpta Vrty” // “Marche / Bo- 
lognola / 14.VII.1937 / Querci / 1400 m” // [blank label] 
// “ZMH 833490”; “correpta Vrty” // “Marche / Bolog- 
nola / 29. VII.1937 / Querci / 1600 m” // [blank label] // 
“ZMH 833491”; “correpta Vrty” // “Marche / Bolognola 
/ 12. VII.1937 / Querci / 1500 m” // [blank label] // “ZMH 
833492”. 

Original locality. Italy: “Bolognola, Central Italy”. 

Remarks. Verity (1920) proposed this name as a race 
of A. hylas (Esper, 1779). According to Kudrna (1983: 
26), this name is unavailable as it denotes a seasonal form 
which is infrasubspecific and hence unavailable. The year 
on the specimen labels (1937) indicate specimens were 
collected 17 years after description (1920) from the same 
location the original holotype material was collected. As a 
result, the black label of the specimens treated as “types” 
is erroneous. The specimen collected from the same geo- 
graphical location from which the type specimen was 
collected is called topotype. According to ICZN (1999), 
a topotype has no formal standing and is not regulated by 
the Code. Therefore, these four specimens are erroneous- 
ly labelled as types, should be annotated as “Not a Type” 
and are treated here as “Non-type” specimens. 


95. olympena Verity, 1936 


Original combination. “L. escheri, Hub. race olympena, 
nom. nov.” Verity, 1936 Ent. Rec. 48: 8. 

Current combination. Polyommatus escheri olympe- 
na (Verity, 1936). 

Current status. Valid subspecies. 

Type material. Syntypes 246 (ZMH 833487-— 
833488) (Fig. 95). “olympena Vrty” // “Macedonia / 
Olympus, 2500’ / July 13, 1935 / Romew” // [blank la- 
bel] // “ZMH 833487”; “olympena Vrty” // “Macedonia 
/ Olympus, 2500’ / July 9, 1936 / Romei” // [blank label] 
/1 “ZMH 833488”. 

Type locality. Greece: “Mt. Olympus, from 850 to 
1500 m.” 

Remarks. Verity (1936) proposed this name as a race 
of L. escheri Hubner, 1823. According to Kudrna (1983: 
34), this race represents an available subspecific name. 


96. ab. privata Schinfeld, 1924 


Original combination. “Lycaena icarus” Rott.-3 “ab. 
privata m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40. 

Current combination. Polyommatus icarus ab. pri- 
vata (Schonfeld, 1924). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827573) (Fig. 96). “14.5. 1921 / KI. Chuchel / Bohmen / 


227 


von Schonfeld” // “Type / Lyc. icarus Rott. 3 / ab. privata 
Schonfeld /1I.E.Z. Guben. 18. Jahr. Nr 6/1924. a.d. nachl. 
v. Schonfeld. / D. Gelpke” // “KI — Chuchel / 14.5.21” / 
“ZMH 827573”. 

Original locality. Czech Republic: “in der Umgebung 
von Prag (bei Kuchelbad)* [in the vicinity of Prague 
(near Kuchelbad)]. 

Remarks. Schoenfeld (1924) proposed this name as 
an aberration of L. icarus Rottemburg, 1775. Therefore, 
as stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


97. nuksani Forster, 1937 


Original combination. “Lycaena candalus nuksani ssp. 
nov.” Forster, 1937 Mitt. Mtinch. Ent. Ges. 27: 62. 

Current combination. Polyommatus (Sublysandra) 
nuksani (Forster, 1937). 

Current status. Valid species. 

Type material. Paratypes 2¢.¢ 29 9, (ZMH 827537-— 
827540) (Fig. 97). “Nord-Ost-Hindukusch / Nuk- 
san-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m 
Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. 
candalus / nuksani Forst. / W. Forster Munchen” // “Coll. 
Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827537”; 
“Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Al- 
penwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. 
Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. 
/ W. Forster Munchen” // “Coll. Bytinski-Salz / Eing. 
Nr. 20 — 60” // “ZMH 827538”; “Nord-Ost-Hindukus- 
ch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500- 
4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type 
/ Lyc. candalus | nuksani Forst. / W. Forster Miinchen” 
// “candalus / nuksani” // “Coll. Bytinski-Salz / Eing. 
Nr. 20 — 60” // “ZMH 827539”; “Nord-Ost-Hindukus- 
ch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500- 
4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type 
/ Lyc. candalus | nuksani Forst. / W. Forster Miinchen” 
// “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 
827540”. 

Type locality. Afghanistan: “Nordost-Hindukusch, 
Nuksan-PaB Nordseite, 3500-4000 m.”. 


98. ab. clorinda Verity, 1938 


Original combination. “Tarucus balkanica, Freyer, with 
I gen. clorinda, nom. nov.” Verity, 1938 Ent. Rec. 50: (8). 

Current combination. Tarucus balkanica ab. clorin- 
da Verity, 1938. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 24'3 (ZMH 
835592-835593) (Fig. 98). “clorinda Vrty” // “Macedo- 
nia / Salonika, 1000’ / May 31, 1935 / Rome” // [blank 
label] // “ZMH 835592”; “clorinda Vrty” // “Macedonia / 


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228 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Lye. arronides - Tr 
afr diac rane ce 


[ZMH -827531| 


2 Paratyeor 
le erdschire 


es 


“Coll Bytinski-Salz 
‘Bing Nr "20, 1960 


avi sssags) © 1 OM 


Plate 13. 85. Phengaris alcon f. lunulata (Warnecke, 1942); 86. Phengaris arionides f. selzeri (Warnecke, 1924); 87. Phengaris 
euphemus peninsulae (Verity, 1923); 88. Cupido argiades ab. depuncta (Hirschke, 1903); 89. Polyommatus (Agrodiaetus) da- 
mon ausonia (Verity, 1913); 90. Polyommatus (Agrodiaetus) pfeifferi (Brandt, 1938), 91. Polyvommatus (Agrodiaetus) alcestis 
(Zerny, 1932); 92. Polyommatus (Agrodiaetus) ardschira (Brandt, 1938); 93. Polyommatus daphnis macra f. squalida Verity, 1920; 
94. Polyommatus dorylas tr. correpta (Verity, 1920). a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


Salonika, 1000’ / May 31, 1935 / Rome1” // [blank label] 
/1 “ZMH 835593”. 

Original locality. Greece: “Salonika”. 

Remarks. Verity (1938) proposed this name as an 
aberration of 7’ balkanica (Freyer, 1844). Therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


99. ahasveros Bytinski-Salz & Brandt, 1937 


Original combination. “Lycaena (Glaucopsyche) pana- 
gaea, H.-Sch. ssp. ahasveros ssp. nov.” Bytinski-Salz & 
Brandt, 1937 Ent. Rec. 49: Suppl. (2). 

Current combination. Turanana panagaea ah- 
asveros (Bytinski-Salz & Brandt, 1937). 

Type material. Paratypes 1432 (ZMH 827556-— 
827569) (Fig. 99). “IRAN: / Keredj (1700 m.) / 6.5.1936 
/ Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / 
ByS+Brdt” // “ssp. ahasveros / ByS + Brdt.” // “Coll. By- 
tinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827556”; “Tran: 
/ Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytins- 
ki-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. 
Nr. 20, 1960” // “ZMH 827557”; “Iran: / Keredj 1700 m 
/ V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratyp- 
oid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827558”; “IRAN: / Keredj (1200 m.) / 6.5.1936 / Brandt” 
// “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827559”; “IRAN: / Keredj (1700 m.) / 6.5.1936 / Brandt” 
// “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827560”; “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / 
coll. Bytinski-Salz” //“Paratypoid” // “Coll. Bytinski-Salz / 
Eing. Nr. 20, 1960” // “ZMH 827561”; “Iran: / Keredj 1700 
m/ V.1936/ leg. F. Brandt / coll. Bytinski-Salz” // “Paraty- 
poid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827562”, “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / 
coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / 
Eing. Nr. 20, 1960” // “ZMH 827563”; “Tran: / Keredj 1700 
m/ V.1936/ leg. F. Brandt / coll. Bytinski-Salz” // “Paraty- 
poid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827564”; “Tran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / 
coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz 
/ Eing. Nr. 20, 1960” // “ZMH 827565”; “IRAN: / Keredj 
(1700 m.) / 10.5.1936 / Brandt” // “Cotypus / L. panagaea 
/ ssp. ahasveros | ByS+Brdt” // “ssp. ahasveros / ByS + 
Brdt.” // “Coll. Bytinski-Salz / Eig. Nr. 20, 1960” // “ZMH 
827566”; “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / 
coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz 
/ Eing. Nr. 20, 1960” // “ZMH 827567”; “Iran: / Keredj 
1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // 
“Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” 
/1 “ZMH 827568”; “X” // “Tran: / Keredj 1700 m / V.1936 / 
leg. F. Brandt / coll. Bytinski-Salz” //“Paratypoid” // “Coll. 
Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827569”. 

Type locality. Iran: Tehran, “Keredj 1700 m.” [Karaj]. 

Current status. Valid subspecies. 


229 


Subfamily Theclinae Swainson, 1831 
Tribe Eumaeini Doubleday, 1847 


100. syra Pfeiffer, 1932 


Original combination. “7hecla ilicis Esp. ssp. n. syra 
m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 1932, Mitt. 
Munch. Ent. Ges. 22: 31. 

Current combination. Satyrium ilicis syra (Pfeiffer, 
1932). 

Current status. Junior subjective synonym of nomi- 
notypical Satyrium ilicis (Esper, 1779). 

Type material. Syntypes 34'\4 (ZMH 827570-827572) 
(Fig. 100). “Syria sept. / amantis / VI.1922” // “Cotypus / 
ilicis / ssp. syra/ Pfeiffer” // “ssp. syra/ Pfeiffer” // “Coll. 
Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 
827570”; “Syria sept. / Taurus / Marasch / 26.V.28 / L. 
Osthelder leg. / 900 m” // “Cotypus / ilicis / ssp. syra / 
Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 
Eing.” // “ZMH 827571”; “Syria sept. / Taurus / Marasch 
/ 26.V.28/L. Osthelder leg. / 1200 m” // “Cotypus / ilicis 
/ ssp. syra/ Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. 
Nr. 20, 1960 Eing.” // “ZMH 827572”. 

Type locality. Turkey: “Syria sept., Taurus, Marasch” 
[Marash]. 


101. brevicaudis Vorbrodt, 1911 


Original combination. “Thecla linceus Esp. (= spini 
Schiff.) brevicaudis Pingeler, 1.1.” Vorbrodt, 1911 Die 
Schmetterlinge der Schweiz. I.: 106. 

Current combination. Satyrium spini brevicaudis 
(Vorbrodt, 1911). 

Current status. Junior subjective synonym of nomi- 
notypical Satyrium spini ({[Denis & Schiffermiuller], 
1775). 

Type material. Syntype | 2 (ZMH 827574) (Fig. 101). 
“Zermatt / Rpe an Rhamnus alp.[ina] / myrmecophil mit 
/ Formica fusca 6.1908 / 8 cl. 28.7.1908 Ping.” // “Co- 
type / v. brevicaudis Ping.” // “Spini var. brevicaudis 
Pung” // “L.29” // “Katalog Nr. / L 2048” // Sig. G. “War- 
necke / Eing. Nr. 5, 1949” // “ZMH 827574”. 

Type locality. Switzerland: “bei Zermatt 1700 m”. 

Remarks. Vorbrodt (1911) proposed this name as a 
form (in the original description and “var.” on the spec- 
imen labels) of “7! Linceus” (sic, recte: [ynceus) (Esper, 
1779). According to article 45.6.4 (ICZN 1999), it is sub- 
specific if first published before 1961 and author express- 
ly used the term “form” or “var”. 


Unpublished, unavailable names 


102. v. romana Hartig 


Original combination. Lycaena cyllarus v. romana 
Hartig. 

Current status. “in litteris’” name and hence not 
available. 


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230 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Original material. Labelled as “Cotype” 3¢.¢ (ZMH 
833472-833474) (Fig. 102). “romana Hartig” // Lazio / 
Formia / 23 4 [19]38 / Querci” // [blank label] // “ZMH 
833472”; “romana Hartig” // Lazio / Formia / 8 4 [19]38 / 
Querci” // [blank label] // “ZMH 833473”; “romana Har- 
tig” // Lazio / Formia / 3 5 [19]38 / Querci” // [blank la- 
bel] // “ZMH 833474”. 

Original locality. Italy: Lazio. 

Remarks. This taxon cannot be found in the literature. 
However, it may refer to Glaucopsyche alexis latina Har- 
tig, 1940 (Mem. Soc. Ent. Ital. 18: 187; Type locality: 
Formia, Latina, Italy). The label data matches the type 
locality of /atina. Papilio cyllarus Rottemburg, 1775 is a 
junior subjective synonym of Papilio alexis Poda, 1761. 


103. ab. albipunctata Hartig 


Original combination. Lysandra coridon ab. albipunc- 
tata Hartig. 

Current status. “in litteris” name and hence not available. 

Original material. Labelled as “Type” 446 (ZMH 
833475833478) (Fig. 103). “Typen” // “19/7 1917 / 
Gottingen / Hohefeld” // “ZMH 833475”; “13/7 1917 / 
Gottingen / Hohefeld” // “ZMH833476”; “18/8 1917 / 
Gottingen / Hohefeld” // “ZMH833477”; “18/7 1917 / 
Gottingen / Hohefeld” // “ZMH 833478”. 

Original locality. Germany: Gottingen. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”) and is hence unavailable. 


Family Nymphalidae Rafinesque, 1815 
Subfamily Danainae Boisduval, 1833 


104. icensis Fuchs, 1954 


Original combination. “Danais gilippus var. icensis” 
Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84. 

Current combination. Danaus gilippus_ icensis 
(Fuchs, 1954). 

Current status. Junior subjective synonym of Danaus 
eresimus erginus (Godman & Salvin, 1897). 

Type material. Lectotype 14 (ZMH 827645), para- 
lectotypes 2346 (ZMH 827643-827644) (Fig. 104). 
“Huayuri / 5/1940 / Type” / “Coll. H. Rodinger / Eing. 
Nr. 30, 1952 / Katalog Nr. L” // “ZMH 827643”; “Ica / 
7/1940 / Type” / “Coll. H. Rodinger / Eing. Nr. 30, 1952 / 
Katalog Nr. L” //“Paralectotype 9 / Danais gilippus / vat. 
icensis / von Fuchs / G. Lamas ‘98” // “Danaus eresimus 
/ erginus Godman + / Salvin 2 / G. Lamas. Det. 1978” // 
“ZMH 827644”; “114 / eresimus. ab / Ica (Pert)” // “Ica / 
7.1928 / Type” // “Coll. H. Rodinger / Eing. Nr. 30, 1952 
/ Katalog Nr. L” // “Lectotype 3 / Danais gilippus / var. 
icensis / von Fuchs / G. Lamas *78” // “Danaus eresimus 
/ erginus Godman + / Salvin ¢ / G. Lamas. Det. 1978” // 
“ZMH 827645”. 


evolsyst.pensoft.net 


Type locality. Peru: “Im Tal von Ica und des Rio 
Grande”. 

Remarks. Fuchs (1954) proposed this name as a “var.” 
of Danais gilippus Cramer, 1775. According to article 
45.6.4 (ICZN 1999), it is subspecific if first published be- 
fore 1961 and author expressly used the term “var.”. 


105. roedingeri Fuchs, 1954 


Original combination. “Danais gilippus var. Rodingeri” 
Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84. 

Current combination. Danaus gilippus roedingeri 
(Fuchs, 1954). 

Current status. Junior subjective synonym of D. g. 
candidus Clark, 1941. 

Type material. Holotype 1 (ZMH 827652), paratypes 
63:2 (ZMH 827649-827651, ZMH 827653-827655) (Fig. 
105). “Danais / gilippus / var. rédingeri & / v. Fuchs” // 
“Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // 
“Std — Peru / Ayacucho — Yucaes / 2500 m 26. 4. 1936.” // 
“Holotype 3 / Danais gilippus / var. Rédingeri / von Fuchs” 
// “Danais gilippus 3 / candidus Clark / G. Lamas. det. 
1978” // “Hbg. — Stid — Peru / Sammelreise 1936. / Eing. 
Nr. 1, 1937.” // “ZMH 827652”; “Ayacucho / 2—3000 m / 
8/1920 / Type” // “D. rédingeri 2” // “Coll. H. Rédinger / 
Eing. Nr. 30, 1952” // “ZMH 827649”; “Type” // “Danais / 
gilippus // var. rédingeri & / v. Fuchs / Rio Jucaes b/ / Aya- 
cucho / 1800-2300” //“Summlung / W. v. Fuchs / Eng. Nr. 
4, 1956” //“ZMH 827650”; “Ayacucho / 2—3000 m/ 8/1920 
/ Type” // “Coll. H. Rédinger / Eing. Nr. 30, 1952 / Katalog 
Nr. L” // “ZMH 827651”; “Ayacucho 28.4” // “Stid — Peru 
/ Oberh. Ayacucho / 29.4.1936.” // “Ayacucho / 2—3000 m 
/ 8/1920 / Type” // “Summlung / W. v. Fuchs / Eing. Nr. 4, 
1956” //“Hbg. — Sud— Peru /Sammelreise 1936. / Eing. Nr. 
1, 1937.” // “ZMH 827653”; “Danais / gilippus / var. r6din- 
geri 2 / v. Fuchs” // “Type” // “Sammlung / W. v. Fuchs / 
Eig. Nr. 4, 1956” // “Std — Peru / Ayacucho — Yucaes / 
2500 m 26. 4. 1936.” // “Hbg. — Sid — Peru / Sammelreise 
1936. / Eing. Nr. 1, 1937.” //“Danais gilippus 9 / candidus 
Clark /G. Lamas. det. 1978” // “Paratype / Danaus gilippus 
9 / var. Rédingeri / von Fuchs” // “ZMH 827654”, “Type” 
// Danais | gilippus / var. rédingeri ° / v. Fuchs / Rio Jucaes 
/ 6/ Ayacucho / 1800—2300” // “ZMH 827655”. 

Type locality. Peru: “... naheren Umgebung von Ay- 
acucho, auf 2000-2600 m” [vicinity of Ayacucho, 2000- 
2600 m]. 

Remarks. Fuchs (1954) proposed this name as a “var.” 
of Danais gilippus (Cramer, 1775). According to article 
45.6.4 (ICZN 1999), it is subspecific if first published be- 
fore 1961 and author expressly used the term “var.”. 


Subfamily Heliconiinae Swainson, 1822 


106. altonevadensis Reisser, 1927 


Original combination. “Argynnis niobe var. nov. altone- 
vadensis” Reisser, 1927 Int. Ent. Z. Guben 20: 373. 


Evolutionary Systematics 5 2021, 193-261 231 


W- chesckel 
4S. 4 


Macedonia 
‘Sal sea = 


935 
tas homme 


4 


Goll. Bytinski-Salz. 
Eing, Nr. 20 - 62. 


oT ay 


Colt, Bytinski-Szl2Coli, 
Eing. Nr. 26, 1960Eing. 


ZMH 827570 


— 0 7 


: 


[ZMH 833473 


[zm 833478 


Plate 14. 95. Polyommatus escheri olympena (Verity, 1936); 96. Polyommatus icarus ab. privata (Schonfeld, 1924); 97. Polyommatus 
(Sublysandra) nuksani (Forster, 1937); 98. Tarucus balkanica ab. clorinda Verity, 1938; 99. Turanana panagaea ahasveros (Bytins- 
ki-Salz & Brandt, 1937); 100. Satyrium ilicis syra (Pfeiffer, 1932), 101. Satyrium spini brevicaudis (Vorbrodt, 1911); 102. Lycaena 
cyllarus v. romana Hartig; 103. Lysandra coridon ab. albipunctata Hartig. a. Dorsal view, b. ventral view. 


1 cm 


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232 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current combination. Fabriciana niobe altone- 
vadensis (Reisser, 1927). 

Current status. Junior subjective synonym of Fabri- 
ciana niobe niobe (Linnaeus, 1758). 

Type material. Syntypes 1 ¢ 192 (ZMH 827656- 
827657) (Fig. 106). “Type # 7 3 / Argynnis niobe var. 
/ altonevadensis Reisser / Int. Ent. Z. guben / Summer 
1927” // “HISPANIA VII 26 / SIERRA NEVADA / O 
BUBACEK” // [blank label] // “ZMH 827656”; “Type # 
4 9 / Argynnis niobe var. / altonevadensis Reisser / Int. 
Ent. Z. guben / Summer 1927” // “HISPANIA VII 26 / 
SIERRA NEVADA / O BUBACEK” // [blank label] // 
“ZMH 827657”. 

Type locality. Spain: “... auf den Hohen der Sierra 
Nevada in Stidspanien ...“. [higher areas of Sierra Neva- 
da in southern Spain]. 

Remarks. Reisser (1927) proposed this name as a “var.” 
of Argynnia niobe (Linnaeus, 1758). According to article 
45.6.4 (ICZN 1999), it is subspecific if first published be- 
fore 1961 and author expressly used the term “var.”. 


(<4 


107. cyrnea Schawerda, 1926 


Original combination. “Argynnis pandora Schiff. var. 
nov. cyrnea” Schawerda, 1926 Z. Ost. Ent. Ver. 11: 109. 

Current combination. Argynnis pandora cyrnea 
Schawerda, 1926. 

Current status. Junior subjective synonym of Argy- 
nnis pandora pandora ({Denis & Schiffermiller], 1775). 

Type material. Syntypes 2¢¢ (ZMH 833447- 
833448) (Fig. 107). “pandora / var. cyrnea Schawerda / 
Cotype [illegible]” // “Evisa Corse / Juli 1921 / Col. O. 
Bubacek” // [blank label] // “ZMH 833447”; “pandora 
/ var. cyrnea Schawerda / Cotype [illegible]” // “Evisa 
Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // 
“ZMH 833448”. 

Type locality. France: “Korsika” [Corsica]. 

Remarks. Schawerda (1926) proposed this name as 
a “var.” of Argynnis pandora ([Denis & Schiffermiuller], 
1775). According to article 45.6.4 ICZN 1999), it is sub- 
specific if first published before 1961 and author express- 
ly used the term “var.”. 


108. argentifasciata Kotzsch, 1937 


Original combination. “Argynnis pandora Schiff. ar- 
gentifasciata ssp. n.” Kotzsch, 1937 Ent. Rdsch. 55: 9. 

Current combination. Argynnis pandora argentifas- 
ciata Kotzsch, 1937. 

Current status. Valid subspecies. 

Type material. Syntypes 24.4 (ZMH 827665-827666) 
(Fig. 108). “West-Hindukusch / Andarab / Gerollzone / 
4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Co- 
typus / A. pandora / ssp. argenti- / fasciata / Kotzsch” 
// “ssp. argentifasciata / Kotzsch” // “ZMH827665”; 
“West-Hindukusch / Andarab / Banu-Ebene / 2000- 


evolsyst.pensoft.net 


2500 m August /leg. H.&E. Kotzsch” //“Cotypus / A. pan- 
dora/ ssp. argenti- / fasciata / Kotzsch” //“ZMH827666”. 

Type locality. Afghanistan: “Andarab im westlichen 
Hindukusch”. 


Subfamily Nymphalinae Rafinesque, 1815 
Tribe Melitaeini Newman, 1870 


109. f. urbanoides Warnecke, 1942 


Original combination. “Melitaea maturna L., 3, n. f. 
urbanoides.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103. 

Current combination. Euphydryas maturna f. ur- 
banoides (Warnecke, 1942). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 24:3 (ZMH 
827658-827659) (Fig. 109). “Holstein / Innien, el. 
/ 1913 Selzer” // “3 nf. urbanoides / Warn. Type / Iris 
1942, 103” // “L.25 / Melitaea maturna - / urbanoides / 
Warn.” // “ZMH 827658”; “Holstein / Innien / [illegible]” 
/1 “VON / G. WARNECKE / W. LINZ. HAMB.” // “Sig. 
W. Linz / Eing. 12 — 74” // “ZMH 827658”. 

Original locality. Germany: “Innien bei Neumutnster 
(Holstein).”; Schleswig-Holstein. 

Remarks. Warnecke (1942) proposed this name as a 
“form” of Melitaea maturna (Linnaeus, 1761). Accord- 
ing to article 45.6.1 (ICZN 1999), it is infrasubspecific if 
the content of the work unambiguously reveals that the 
name was proposed for an infrasubspecific entity, and 
is hence unavailable. The title already indicates that this 
form constitutes an individual aberration and thus an un- 
available name. 


110. tenuicula Verity, 1919 


Original combination. “Melitaea athalia race tenuicula 
mihi” Verity, 1919 Ent. Rec. 31: 193-194. 

Current combination. Melitaea athalia tenuicula 
Verity, 1919. 

Current status. Valid subspecies. 

Original material. Labelled as “Type” 64° (ZMH 
833455—-833460) (Fig. 110). “tenuicula Vrty” // helvet- 
ica Ruhl” // “Marche / Bolognola / 28 VI 1937 / Quer- 
ci.” // [blank label] // “ZMH 833455”; “tenuicula Vrty” 
// helvetica Ruhl” // “Marche / Bolognola / 15 VI 1937 
/ Querci.” // [blank label] // “ZMH 833456”; “tenuicula 
Vrty” // “Marche / Bolognola / 17 VI 1937 / Querci.” // 
[blank label] // “ZMH 833457”; “tenuicula Vrty” // hel- 
vetica Ruhl” // “Marche / Bolognola / 18 VI 1937 / Quer- 
ci.” // [blank label] // “ZMH 833458”; “tenuicula Vrty” // 
“Marche / Bolognola / 28 VI 1937 / Querci.” // [blank la- 
bel] // “ZMH 833459”; “tenuicula Vrty” // “Marche / Bo- 
lognola / 27 VI 1937 / Querci.” // [blank label] // “ZMH 
833460”. 

Original locality. Italy: “Bolognola in the Sibillini 
Mountains, 1200 m”. 


Evolutionary Systematics 5 2021, 193-261 233 


: Coll. H,.Rédinger 
: Eing. Nr. 30, 1952 
‘Katalog Nr. L 


Se 
Danaus eresim us 3 
Jabvin 
G.Lamas. det. 1976 


Z7MH_827645| 


Yerrscr¢ i 


haa: 9¢Lf ee 
ohn vedenaered 
Ks ar 


hace ‘ae 
Danaiy 9 lippus 
yer. pack wey 


G. Lamas. det. 1978 


F ~~ Sammlung a 
W. v. Fuchs : 


Evisa_ Corse 
Jul 1921 
Col.O,.Bubacek 


= 


HISPANIA VII 26 
SIERRA. NEVADA 
Q BUBACEK 


a 


ZMH_827656 


107b- 


West-Hindukusch 
@ Andarab 
Geridlzone 
4000-4500m Mitte August , 
leg. H.&E.Kotzsch 


[ZMH 827665 


IZMH 827658| 


11cm 


Plate 15. 104. Danaus gilippus icensis (Fuchs, 1954); 105. Danaus gilippus roedingeri (Fuchs, 1954); 106. Fabriciana niobe alto- 
nevadensis (Reisser, 1927); 107. Argynnis pandora cyrnea Schawerda, 1926; 108. Argynnis pandora argentifasciata Kotzsch, 1937; 
109. Euphydryas maturna f. urbanoides (Warnecke, 1942). a. Dorsal view, b. ventral view. 


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234 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Remarks. Verity (1919) proposed this name as a “race” 
of Melitaea athalia (Rottemburg, 1775). According to 
Kudrna (1983: 83-84), this name is available as subspecif- 
ic. The year on the specimen labels (1937) indicate spec- 
imens were collected 18 years after description (1919) 
from the same location the original holotype material was 
collected. As a result, the black label of the specimens 
treated as “types” is erroneous. The specimen collected 
from the same geographical location from which the type 
specimen was collected is called topotype. According to 
ICZN (1999), a topotype has no formal standing and 1s not 
regulated by the Code. Therefore, these six specimens are 
erroneously labelled as types, should be annotated as “Not 
a Type” and are treated here as “Non-type” specimens 


111. varissima Verity, 1914 


Original combination. “MVelitaea [parthenie Bkh] varia 
Meyer-Dur varissima nom. nov.” Verity, 1914 (1913) 
Boll. Soc. Ent. Ital. 45: 210. 

Current combination. Melitaea varia varissima Ver- 
ity, 1914. 

Current status. Valid subspecies. 

Original material. Labelled as “Type” 63'¢ (ZMH 
833461—833466) (Fig. 111). “varissima Vrty” // “varia 
Meyer-Dir” // “Marche / Bolognola / 25.VI.1937 / Quer- 
ci” // [blank label] // “ZMH 833461”; “varissima Vrty” // 
“Marche / Bolognola / 20. V1.1937 / Querci” // [blank label] 
/!“ZMH 833462”; “varissima Vrty” // “Marche / Bologno- 
la/14.V1.1937 / Querci” // [blank label] // “ZMH 833462”; 
“yarissima Vrty” // “Marche / Bolognola / 14.VI.1937 
/ Querci” // [blank label] // “ZMH 833463”; “varissi- 
ma Vrty” // “Marche / Bolognola / 5.VI.1937 / Querci” 
// [blank label] // “ZMH 833464”; “varissima Vrty” // 
“Marche / Bolognola / 22. V1.1937 / Querci” // [blank label] 
/!“ZMH 833465”; “varissima Vrty” // “Marche / Bologno- 
la /25.V1.1937 / Querci” // [blank label] // “ZMH 833466”. 

Original locality. Italy: Bologna “Nei Faggeti di 
Bologna, nella localita Costarotara e a Piano Astore dei 
Monti Sibillini, da 1200 a 1600 m. d’altitudine” [Bolo- 
gna, vicinity of Costarotara and Piano Astore of the Sibil- 
lini Mountains, 1200-1600 m.]. 

Remarks. Verity (1919) proposed this name as a “race” 
of Melitaea varia Meyer-Dir, 1851. According to Kudrna 
(1983: 84), this name is available as subspecific. The year 
on the specimen labels (1937) indicate specimens were 
collected 23 years after description (1914) from the same 
location the original holotype material was collected. As a 
result, the black label of the specimens treated as “types” 
is erroneous. The specimen collected from the same geo- 
graphical location from which the type specimen was col- 
lected is called topotype. According to ICZN (1999), a 
topotype has no formal standing and is not regulated by 
the Code. Therefore, these six specimens are erroneously 
labelled as types, should be annotated as “Not a Type” 
and are treated here as “Non-type” specimens. 


evolsyst.pensoft.net 


112. riloensis Ziillich, 1936 


Original combination. “Melitaea cinxia L. var. riloensis 
Ziillich.” Zullich, 1936 Z. Ost. Ent. Ver. 21: 27. 

Current combination. Melitaea cinxia riloensis Ziil- 
lich, 1936. 

Current status. Junior subjective synonym of Meli- 
taea cinxia Clarissa Staudinger. 

Type material. Syntypes 2¢¢ (ZMH 827646- 
827647) (Fig. 112). “COTYPE / Mel. cinxia L. / 3 v. ri- 
loensis / Z. d. O. Ent. V. 1936 / DR. ZULLICH, WIEN” 
// “BULGARIEN JUNI 1923 / RYLA GEB. 1200 bis 
1800 m/O. BUBACEK WIEN” // [blank label] // “ZMH 
827646”; “COTYPE / Mel. cinxia L. / 3 v. riloensis / Z. 
d. O. Ent. V. 1936 / DR. ZULLICH, WIEN” // “BUL- 
GARIEN JUNI 1923 / RYLA GEB. 1200 bis 1800 m / 
O. BUBACEK WIEN” // [blank label] // “ZMH 827647”. 

Type locality. Bulgaria: “auf dem Steilhang ostlich 
des Riloklosters in einer Hohe von etwa 1200 m.” [at 
a steep slope east of the Rilo monastry at an altitude of 
about 1200 m]. 1901. 

Remarks. Zillich (1936) proposed this name as a 
“var.” of M. cinxia (Linnaeus, 1758). According to arti- 
cle 45.6.4 (ICZN 1999), it is subspecific if first published 
before 1961 and author expressly used the term “var.”. 


113. ab. mediofasciata Bubacek, 1926 


Original combination. “Melitaea dejone H.G. ab. medi- 
ofasciata Bub. ab. nova.” Bubacek, 1926 Verh. Zool.-Bot. 
Ges. Wien 74—75: (9). 

Current combination. Melitaea deione ab. medio- 
fasciata Bubacek, 1926. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
833454) (Fig. 113). “Type / Mel. rondoui / ab. mediofas- 
ciata / Otto Bubacek” // “France, Gedre / Hautes Pyrenées 
/ Juli 1923 / Col. Otto Bubacek” // [blank label] // “ZMH 
833454”. 

Original locality. France: Pyrenées. 

Remarks. Bubacek (1926) proposed this name as an 
aberration of M. dejone Geyer, 1832. As stated by article 
45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific 
name (the author used “aberration”, “ab.”) and is hence 
unavailable. 


114. ab. johni Stauder, 1922 


Original combination. “M/. dictynna ab. johni” Stauder, 
1922 Ent. Anz. 2: 44. 

Current combination. Melitaea diamina ab. johni 
Stauder, 1922. 

Current status. Infrasubspecific and hence unavail- 
able name. 


Evolutionary Systematics 5 2021, 193-261 


Original material. Labelled as “Type” 14 (ZMH 
827648) (Fig. 114). “Lainau 800 m / [illegible] / 1.VII.17 
Schawerda” // “Type” // “dictynna f. / johni Stauder” // 
“Type” // “ZMH 827648”. 

Original locality. Austria: “Salzkammergut (Lainau- 
tale, Hallstatt)”. 

Remarks. Stauder (1922) proposed this name as an 
aberration of M. dictynna Esper, 1778. As stated by arti- 
cle 45.6.2 (ICZN 1999) it is deemed to be an infrasubspe- 
cific name (the author used “aberration”, “ab.”) and is 
hence unavailable. 


115. balcanica Ziillich, 1936 


Original combination. “MVelitaea trivia Schiff. var. bal- 
canica Zullich.” Zullich, 1936 Z. Ost. Ent. Ver. 21: 27. 

Current combination. Melitaea trivia balcanica 
Ziillich, 1936. 

Current status. Junior subjective synonym of Meli- 
taea trivia lathon Fruhstorfer, 1917. 

Type material. Syntypes 246 (ZMH 827663- 
827664) (Fig. 115). “BULGARIEN JUNI 1928 / RYLA 
GEB. 1200 bis 1800 m/O. BUBACEKR” // [blank label] // 
“COTYPE / Mel. trivia/ Gv. balcanica / Z. d. O. Ent. V. 
1936 /DR. ZULLICH, WIEN” // “ZMH 827663”; “BUL- 
GARIEN JUNI 1928 /RYLA GEB. 1200 bis 1800 m/O. 
BUBACEK” // [blank label] // “COTYPE / Mel. trivia / 
3 v. balcanica / Z. d. O. Ent. V. 1936 / DR. ZULLICH, 
WIEN” // “ZMH 827664”. 

Type locality. Bulgaria: “in den gebirgigen Teilen der 
zentralen und 6stlichen Balkanhalbinsel (Krain, Bosnien, 
Herzegowina etc.)”. [in the mountainous parts of the cen- 
tral and eastern Balkan peninsula]. 

Remarks. Zillich (1936) proposed this name as a va- 
riety of M. trivia ([Denis & Schiffermitller], 1775). Ac- 
cording to article 45.6.4 (ICZN 1999), it is subspecific if 
first published before 1961 and author expressly used the 
term “var.”. 


116. catapelioides Stauder, 1918 


Original combination. “Velitaea trivia catapelioides nov. 
subspec. m.” Stauder, 1918 Zeit. Wiss. Ins. Biol. 14: 57. 
Current combination. Melitaea trivia catapelioides 
Stauder, 1918. 
Current status. Junior subjective synonym of nomino- 
typical Melitaea trivia ({[Denis & Schiffermtller], 1775). 
Original material. Labelled as “Type” 14 (ZMH 
827662) (Fig. 116). “Type” // “Luen / Calabria mer. / 
Aspromonte 950 m. / 6.VII 1920 / H. Stauder legit.” // 
“trivia catape - / lioides Stauder” // “Type” // “Sig. G. 
Warnecke / Eing. Nr. 5, 1945” // “ZMH 827662”. 
Original locality. Italy: “Unteritalien (Kalabrien)”. 
Remarks. The year on the specimen label (1920) 
indicates specimen was collected two years after 


230 


description (1918) from the same location (..e., Italy, 
Kalabrien) the original holotype material was collected. 
As a result, the label of the specimen treated as “Type” 
is erroneous. The specimen collected from the same 
geographical location from which the type specimen was 
collected is called topotype. According to ICZN (1999), 
a topotype has no formal standing and is not regulated 
by the Code. Therefore, this specimen is erroneously 
labelled as type, should be annotated as “Not a Type” and 
is treated here as “Non-type” specimen. 


117. ignasiti de Sagarra, 1926 


Original combination. “Melitaea trivia Schiff., rassa 
ignasiti, nova rassa” de Sagarra, 1926 Butl. Inst. Catal. 
Hist. Nat. (2)6: 131. 

Current combination. Melitaea trivia ignasiti de 
Sagarra, 1926. 

Current status. Valid subspecies. 

Type material. Paralectotypes 1419 (ZMH 827660-— 
827661) (Fig. 117). “CATALONIA / Seva 700 m. / 3.8.25 
Romei” // “M. trivia — ignasiti Sag. / co-type 4 Querci” 
/! “ZMH 827660”; “CATALONIA / Seva 700 m. / 3.8.25 
Romei” // “Sammlung / R. Jahnig / Eing. Nr.29,1949” // 
“M. trivia — ignasiti Sag. / co-type 2 Querci” // “L.28” // 
“ZMH 827661”. 

Type locality. Spain: Catalonia, “Martorell”. 

Remarks. de Sagarra (1926) proposed this name as 
a “rassa = race” of M. trivia ([Denis & Schiffermullar], 
1775). The term “rassa = race” which was employed by 
de Sagarra in 1926 was often used in the current sense 
of subspecies. As a result, the first line of article 45.6 
(ICZN 1999) — the rank denoted by a species-group 
name following a binomen is subspecific — is applied 
here, it is thus available as subspecific. Higgins (1941) 
synonymised it with M. trivia pseudodidyma Rebel, 
1905 but he reconsidered it as a valid subspecies in his 
subsequent books. Garcia-Barros et al. (2013) made 
a comment in Fauna Ibérica that “the orange color of 
the Iberian specimens is lighter than those from other 
European areas; those have been attributed to subspecies 
“ignasiti de Sagarra, 1926”, which, however, does not 
seem significantly different. Furthermore, it does not 
seem significantly differentiated from a genetic point of 
view (Wahlberg pers. comm.).” The taxon ignasiti was 
reinstated as a valid species by van Oorschot and Coutsis 
(2014), and this was accepted by Macia et al. (2015) 
who designated lectotype and paralectotypes. Therefore, 
specimens at ZMH are by default paralectotypes. 
However, most other recent authors (e.g., Kudrna 2019) 
treat this taxon as a synonym to M. trivia, because 
there is no evidence for species status. Even Oorschot 
and Coutsis (2014) mention intermediates indicating 
hybridisation. It should be noted that the species name is 
not mentioned in the latest European checklist (Wiemers 
et al. 2018). 


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236 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


BS . ) ee I ub aia 4 a rr 
Leinwicate \Ivta AMA Mitek Nin iy 
ea A. <7 GP he Belated gaan ash 

Marche 
Bolognola 
LE wyP1987 


uercl, 


Wdvahue Ral 
ee 


wlth hetediact. 
..... Ab, medtolasciaty. 
7] a acche 


COTYPE 
Me aural, 


France, Gédre 
Hautes Pyrenées 
Juli 1923 
Col. Otto Bubacek 


Tu, uloonsts 
Z.4.b6-EntV. 1936 


DR. ZULLICH, WIEN 


BULGARIEN JUN] 1523 
RYLA GES. 1200 bes 1800 m 
Q. BUBACEK WIEN 


[ZMH 833454! 


BULGARIEN ..JUNI 1928 
RYLA GEB. 1200 bis 1800 m 
0. BUBACEK WIEN 


COTYPE 
Md. tuvia 
Ov. baleanica 
2ST Ent Ve 1936 


DR. ZOLIICH, WIEN 


ZMH 827663 


CATALONIA 
Seva 700 m. 
4.8.25 Romei 


Calabria mer. 


Asprofaonte m, 
OV77 Leas 
H. Stauder legit. 


[ZMH _827660| 


Sig. G. Warnacke 
Elng. Nr.5 ,1948 


ZMH _ 827662 


Plate 16. 110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis 
Ziullich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea 
trivia balcanica Zillich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. 
a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 
Tribe Nymphalini Rafinesque, 1815 


118. guhni Tschauner, 1926 


Original combination. “Vanessa urticae var. guhni” 
Tschauner, 1926 Int. Ent. Z. 20: 229, 273. 

Current combination. Aglais  urticae 
(Tschauner, 1926). 

Current status. Junior subjective synonym of Ag/ais 
urticae urticae (Linnaeus, 1758). 

Type material. Paratype 1 (ZMH 827667) (Fig. 118). 
“Paratypoid” // “Vanessa urticae / nov. var. guhni / Para- 
type e. coll. / Tschauner, Berlin 1926” // “Coll. Bytins- 
ki-Salz / Eing. Nr. 20, 1960” // “ZMH 827667”. 

Type locality. Germany: Berlin. 

Remarks. This case seems ambiguous, because 
Tschauner (1926) writes that the new name constitutes a 
local variation, if not a new race, although he also notes 
that he is not sure whether the variation is really heritable. 
On page 273 he names it again as “Vanessa urticae var. 
Guhni nov. spec.“ [sic!]. See also the critical remark by 
Kitt (1927). 


guhni 


119. f. clarior Warnecke, 1959 


Original combination. “Araschnia levana (Linnaeus) 
gen. vern. n. f. clarior’ Warnecke, 1959 Mitt. Faun. Arb. 
Gem. Schlesw. Holst. 1 (10.): 53. 

Current combination. Araschnia levana f. clarior 
Warnecke, 1959. 

Current status. Infrasubspecific and thus unavail- 
able name. 

Original material. Labelled as “Type” 26’ (ZMH 
825009-825010) (Fig. 119). “Schleswig / Elsdorfer Sch. 
/ 20.3.97” // “n.f. clarior W. / Coll. Warnecke / Eing. Nr. 
5-1949” // “ZMH 825009”; “Schleswig / Elsdorfer Sch. 
/c. 1. 20.3.97” // “nf. clarior W. / Coll. Warnecke / Eing. 
Nr. 5-1949” // “ZMH 825010”. 

Original locality. Germany: Schleswig-Holstein. 

Remarks. Warnecke (1959) proposed this name as an 
aberration of A. /evana (Linnaeus, 1758). According to 
article 45.6.1 (ICZN 1999), it is infrasubspecific if the 
content of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity, and is hence 
unavailable. From the text of the description it is unam- 
biguously clear that the author is describing an aberration 
and not a subspecies. 


120. ab. marginelineata Horch, 1932 


Original combination. “Araschnia levana L. gen. aest. 
prorsaab. marginelineata” Horch, 1932 Int. Ent. Z. 26:338. 
Current combination. Araschnia levana prorsa ab. 
marginelineata Horch, 1932. 
Current status. Infrasubspecific and hence unavail- 
able name. 


237 


Original material. Labelled as “Kotype” 14 (ZMH 
825011) (Fig. 120). “f. marginelineata / Horch / Gub. I. E 
Z; 26., 1932/3, / P. 337.” // “Umgeg. v. Hamburg / Rad- 
bruch / e. |. 10.7.32 / R. Horch Hamburg” // “Kotype” // 
“ZMH 825011”. 

Original locality. “Luneburger Heide”, Hamburg 
[Germany]. 

Remarks. Horch (1932) proposed this name as an 
abberation of A. /evana (Linnaeus, 1758). This species 
is known for having two forms, /evana and prorsa that 
represent the spring and summer broods respectively. As 
stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


Subfamily Satyrinae Boisduval, 1833 
Tribe Satyrini Boisduval, 1833 


121. alphea Warnecke, 1919 


Original combination. “Satyrus arethusa Schiff. nov. 
subsp. alphea Warn.” Warnecke, 1919 Int. Ent. Z. 
12: 182. 

Current combination. Arethusana arethusa alphea 
(Warnecke, 1919). 

Current status. Junior subjective synonym of Arethu- 
sana arethusa arethusa ({Denis & Schiffermiuller], 1775). 

Type material. Syntype 13 (ZMH 827676) (Fig. 121). 
“Type” // nov. v. Alphea” / Warn.” // “Type” // “Griechen- 
Id.” // “ZMH 827676”. 

Type locality. Greece: “Griechenland, Velucci und 
Parnass”’. 


122. asperomontana Stauder, 1921 


Original combination. “Satyrus circe L. subsp. n. asper- 
omontana” Stauder, 1921 Dt. Ent. Z. Iris 35: 28. 

Current combination. Brintesia circe asperomon- 
tana (Stauder, 1921). 

Current status. Junior subjective synonym of Brinte- 
sia circe circe (Fabricius, 1775). 

Type material. Syntype 13 (ZMH 827680) (Fig. 122). 
“1/7 / Calabria mer. / Asperomonte 1200 m. / Polsi 1920 
/ H. Stauder legit.” // “Type” // “circe / asperomontana / 
Type” // “ZMH 827680”. 

Type locality. Italy: “Polsibeken, Aspromonte 1350- 
1500 m.” 


123. f. rufobrunnea Warnecke, 1942 


Original combination. “Coenonympha arcania L. n. f. 
rufobrunnea.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102. 
Current combination. Coenonympha arcania f. ru- 
fobrunnea Warnecke, 1942. 
Current status. Infrasubspecific and hence unavail- 
able name. 


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238 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


SAL, a1’; 
a erin VY 
uv e ‘ ty! 


mia ~ 
iy ae 


ae See be 
Coli. Pytinski-Saiz ; 
ZMH 827667 


' Coli. Warnecke 
fing. Nr §-1948 


f | Vs aa Of uta le 


Kg ae 46. 14343 
ft 992 ¢ 


Umgeg.v-Hamburg 
bruce 


Rad h 
C4732, 


R.Horch Hamburg: 


ZMH 827680 


1cm 


Plate 17. 118. Aglais urticae guhni (Tschauner, 1926); 119. Araschnia levana f. clarior Warnecke, 1959; 120. Araschnia levana 
prorsa ab. marginelineata Horch, 1932; 121. Arethusana arethusa alphea (Warnecke, 1919); 122. Brintesia circe asperomontana 
(Stauder, 1921). a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


Original material. Labelled as “Type” 1¢ (ZMH 
827677) (Fig. 123). “Itzehoe / (Holstein)” // “n.f. rufobrun- 
nea Warn. / Type / Iris 1942, p. 102” // “Katalog Nr. 519 / 
HL” // “Coenon. arcania - / f. rufobrunnea / Warn. / L.38” // 
“Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827677”. 

Original locality. Germany: Schleswig-Holstein, 
Itzehoe. 

Remarks. Warnecke (1942) proposed this name as a 
form of C. arcania (Linnaeus, 1761). According to article 
45.6.1 ICZN 1999), it is infrasubspecific if the content of 
the work unambiguously reveals that the name was pro- 
posed for an infrasubspecific entity, and is hence unavail- 
able. From the text it is clear that the author describes an 
individual form / aberration. 


124. tyrrhena Stauder, 1915 


Original combination. “Coenonympha arcania tyrrhena 
m. subsp. nov.” Stauder, 1915 Zeit. Wiss. Ins. Biol. 11: 1, 
is aed MS Si 

Current combination. Coenonympha arcania tyr- 
rhena Stauder, 1915. 

Current status. Junior subjective synonym of Co- 
enonympha arcania arcania (Linnaeus, 1761). 

Type material. Syntype 13 (ZMH 827761) (Fig. 124). 
“Type” // “Calabria / Umg. Paola-Co-/Senza 18.6.1920 / 
H. Stauder” // “Type / v. tyrrhena Stauder” // “arcania v. 
/ tyrrhena Staud.” // “ZMH 827761”. 

Type locality. Italy: Monte Martinello bei 1000 m. 
[Castello Cabiaglio, Province of Varese, Italy]. 


125. ab. energica Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova energica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. 
Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
energica Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 1¢ (ZMH 
827740) (Fig. 125). “Type / Coen. corinna Hb. / ab. energi- 
ca Bub. / Otto Bubacek Z.B.G. 1922” //“Evisa Corse / Juni 
1921 / Col. O. Bubacek” // [blank label] // “ZMH 827740”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.”) 
and is hence unavailable. 


126. ab. caeca Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova caeca Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. 
Wien 72: (30). 


239 


Current combination. Coenonympha corinna ab. 
caeca Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827741) (Fig. 126). “Type / Coen. corinna Hb. / ab. cae- 
ca Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / 
Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 
827741”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.’’) 
and is hence unavailable. 


127. ab. anophthalmica Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova anophthalmica Bub.” Bubacek, 1923 Verh. Zool. 
Bot. Ges. Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
anophthalmica Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827742) (Fig. 127). “Type / Coen. corinna Hb. / ab. 
anophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // 
“Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank la- 
bel] // “ZMH 827742”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.’) 
and is hence unavailable. 


128. ab. saturata Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova saturata Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. 
Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
saturata Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827743) (Fig. 128). “Type / Coen. corinna Hb. / ab. 
saturata Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa 
Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // 
“ZMH 827743”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.’’) 
and is hence unavailable. 


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240 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Vip 
Pry Be, fr COL 


bs ee 


Baaeneste SNA 3276771 


Calabria . 
Umg.Padla-Co- 


senza fi 6, 19 
2 TeStauder=? 


V, Cgrrhtica Maiadip 


Lype 


Ab.caeeq Bub... 
to Buhacek.ZB.os gAL, 


Evisa Corse 
| Juli 1921 
Col.0.Bubacek 


Evisa. Corse 
Junl iyza 
Col.O.Bubacek 


Pe 


Evisa, Corse 
Jull 1921 
Col.O.Bubacek 


Evisa: Corse " 
Jum 1921 
Col.0.Bubacek 


= 


ZMH 827743 


Ly pe 
Coen,corenna Hp. 
| ad neorfteans Bub, 
Oka Bibixcel 2 BO/444, 


| ZMH 827744 
Col.O.Bubacek 


ZMH 827742 


Plate 18. 123. Coenonympha arcania f. rufobrunnea Warnecke, 1942; 124. Coenonympha arcania tyrrhena Stauder, 1915; 
125. Coenonympha corinna ab. energica Bubacek, 1923; 126. Coenonympha corinna ab. caeca Bubacek, 1923; 127. Coenonympha 
corinna ab. anophthalmica Bubacek, 1923; 128. Coenonympha corinna ab. saturata Bubacek, 1923; 129. Coenonympha corinna ab. 
nigricans Bubacek, 1923. a. Dorsal view, b. ventral view. 


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Evolutionary Systematics 5 2021, 193-261 


129. ab. nigricans Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova nigricans Bub.” Bubacek, 1923 Verh. Zool. Bot. 
Ges. Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
nigricans Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827744) (Fig. 129). “Type / Coen. corinna Hb. / ab. nig- 
ricans Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse 
/ Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 
827744”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.”) 
and is hence unavailable. 


130. ab. macrophthalmica Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova macrophthalmica Bub.” Bubacek, 1923 Verh. Zool. 
Bot. Ges. Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
macrophthalmica Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 24'3' (ZMH 
827745—827746) (Fig. 130). “Type / Coen. corinna Hb. / 
ab. macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” 
// “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank 
label] // “ZMH 827745”; “Type / Coen. corinna Hb. / ab. 
macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // 
“Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank la- 
bel] // “ZMH 827746”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.”) 
and is hence unavailable. 


131. ab. gynandra Bubacek, 1923 


Original combination. “Coenonympha corinna Hb. ab. 
nova gynandra Bub.” Bubacek, 1923 Verh. Zool. Bot. 
Ges. Wien 72: (30). 

Current combination. Coenonympha corinna ab. 
gynandra Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 246 (ZMH 
827747—827748) (Fig. 131). “Type / Coen. corinna Hb. 
/ ab. gynandra Bub. / Otto Bubacek Z.B.G. 1922” // 


241 


“Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank 
label] // “ZMH 827747”; “Type / Coen. corinna Hb. / ab. 
gynandra Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa 
Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // 
“ZMH 827748”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of C. corinna (Hubner, 1804). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.’’) 
and is hence unavailable. 


132. f. pulcherrima Warnecke, 1942 


Original combination. “Coenonympha iphis Schiff., 9 
n.f. pulcherrima.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 
102. 

Current combination. Coenonympha glycerion f. 
pulcherrima Warnecke, 1942. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 12 (ZMH 
827738) (Fig. 132). “Molln 1. lotz / 30.6.35 [illegible]” 
/! “30.6.35 / Molln” // “9 n.f. pulcherrima / Warn. Type / 
Tris 1942, 102” // “Coll. Warnecke / Eing. Nr. 5, 1949” // 
“Type” // “ZMH 827738”. 

Original locality. Germany: “Molln (Lauenburg)*; 
Schleswig-Holstein, Herzogtum Lauenburg, MOlIn. 

Remarks. Warnecke (1942) proposed this name as 
a form of C. iphis ([Denis & Schiffermuller], 1775). 
According to article 45.6.1 (ICZN 1999), it is infra- 
subspecific if the content of the work unambiguously 
reveals that the name was proposed for an infrasubspe- 
cific entity, and is hence unavailable. From the text it 
is clear that the author describes an aberration and not 
a subspecies. 


133. ab. oikeia Dannehl, 1927 


Original combination. “Coenonympha iphis Schiff. ab. 
oikeia Dhl.” Dannehl, 1927 Mitt. Minch. Ent. Ges. 17: 5. 

Current combination. Coenonympha glycerion ab. 
oikeia Dannehl, 1927. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
835591) (Fig. 133). “25.6” // “Italia central. / Mti. Sim- 
bruini/ coll. F Dannehl” // “oikeia / Dhl. / Cotype. / G.” // 
“Cotype” // “25.6. 1926 / Mti. Simbruini / Mittel.Italien” 
// “ZMH 835591”. 

Original locality. Italy: “Mt. Simbruini von etwa 
1400 m Hohe”. 

Remarks. Dannehl (1927) proposed this name as an 
aberration of C. iphis ({Denis & Schiffermiller], 1775). 
As stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


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242 


134. iranica Schwingenschuss, 1939 


Original combination. “Coenonympha leander Esp. 
nov. ssp. iranica Sch.” Schwingenschuss, 1939 Ent. Z. 
53: 62: 

Current combination. Coenonympha leander irani- 
ca Schwingenschuss, 1939. 

Current status. Valid subspecies. 

Type material. Syntype 14 (ZMH _ 827714) 
(Fig. 134). “Pers. Elbursgeb. / Kendevanpass / 2800 m, 
3.-8.7.36 / Schwingenschuss” // “Cotypus” // “ssp. irani- 
ca Schwing.” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” 
/1 “ZMH 827714”. 

Type locality. Iran: “Am Kendevan (Sch., W., Pf.) von 
2600 m bis 3000 m” [Kandovan pass]. 


135. rhenana Gradl, 1933 


Original combination. “Coenonympha oedipus F. sub- 
species rhenana” Gradl, 1933 Int. Ent. Z. 27: 261. 

Current combination. Coenonympha_ oedippus 
rhenana Gradl, 1933. 

Current status. Junior subjective synonym of nomi- 
notypical Coenonympha oedippus (Fabricius, 1787). 

Type material. Syntypes 4¢¢ (ZMH 827722- 
827725) (Fig. 135). “Feldk / Vorarlberg / Prof. F. Gradl 
/ 31.7.32 / Rheinau” // “Subsp. / rhenana / F-Gradl” // 
“Cotype” // “L.23 / Coenonympha / oedippus - | rhena- 
na Gradl” // “Sig. G. Warnecke / Eing Nr 5, 1949” // 
“ZMH 827722”; “Vorarlberg / Prof. F. Gradl / Rheinauen 
/12.7.27 // “Subsp. / rhenana / F-Gradl” // “Cotype” // 
“ZMH 827723”; “Vorarlberg / Prof. F. Gradl / Feldkirch. 
Rhein / auen. 12.7.31” // “Subsp. / rhenana / F-Gradl” 
// “Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827724”; “Vorarlberg / Prof. F. Gradl / 
Feldk-Rheinau / 28.7.32” // “Subsp. / rhenana / F-Gradl” 
/1 “Co-Type” // “Type” // “ZMH 827725”. 

Type locality. Germany: “in den Niederungen des 
Rheintales”. [in the lower region of the Rhine valley]. 


136. galvagnii Stauder, 1923 


Original combination. “C. p. galvagnii m. nova sub- 
spec. (partim aberr.)” Stauder, 1923 Zeit. Wiss. Ins. Biol. 
1259 ty Ser a6 13: 

Current combination. Coenonympha pamphilus 
galvagnii Stauder, 1923. 

Current status. Junior subjective synonym of nomi- 
notypical Coenonympha pamphilus (Linnaeus, 1758). 

Type material. Syntype 14 (ZMH 827739) (Fig. 136). 
“Type” / “H. Stauder / Litorale austr. / Triest Umgbg. / 
12/8/1913” // “pamphilus | galvagnii Staud.” // “Type” // 
“Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827739”. 

Type locality. Italy: Bologna “Bei Grado auf den La- 
gunen in der g. v. und sonst im Kustengebiete, bei Triest, 
Gorz.” [Bologna, vicinity of Costarotara and Piano As- 
tore of the Sibillini Mountains, 1200-1600]. 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


137. f. maculata Warnecke, 1942 


Original combination. “Coenonympha tiphon Rott., °, 
n. f. maculata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103. 

Current combination. Coenonympha tullia f. macu- 
lata Warnecke, 1942. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 229 (ZMH 
827762-827763) (Fig. 137). “Bahrenfeld / 1913” // “9 
n.f. maculata / Warn. Cotype / Iris 1942, 103” // “Coll. 
Warnecke / Eing. Nr. 5, 1949” // “ZMH 827762”; “Sege- 
berg / Juli 13” // “Coll. Warnecke / Eing. Nr. 5, 1949” // 
“ZMH 827763”. 

Original locality. Unclear; Germany: Bahrenfeld, 
Hamburg “aus Holstein.”; publication states: Holstein, la- 
bel of cotype specimen from Warnecke states Bahrenfeld, 
?, (one further specimen without type label: Segeberg) 

Remarks. Warnecke (1942) proposed this name as 
a form of C. tiphon (Rottemburg, 1775). According to 
article 45.6.1 (ICZN 1999), it is infrasubspecific if the 
content of the work unambiguously reveals that the name 
was proposed for an infrasubspecific entity, and 1s hence 
unavailable. From the text and already from the title of 
the paper it is clear that the author describes an aberra- 
tion. Papilio tiphon Rottemburg, 1775 is junior subjec- 
tive synonym of Coenonympha tullia (Miller, 1764). 


138. flavofasciata Heyne, 1895 


Original combination. “Evebia flavofasciata Heyne.” 
Heyne, [1895]; in Ruhl & Heyne, Die Pal. Gross-Schmett. 
1: 805. 

Current combination. Erebia flavofasciata Heyne, 
1895. 

Current status. Valid species. 

Type material. Paratype 14 (ZMH 833469) (Fig. 138). 
“Campolungo / Pass / Oberst v. Nolte” // “Flavofasciata” 
// “Type oder Paratype / e coll. von Nolte / s. Heyne-Ruhl, 
1895, 805, / der keine Samml. hatte. / von Fuchs-Hbg 
ded.” // “ZMH 833469”. 

Type locality. Italy: Tessin, Campolungo pass close 
to Fusio. 


139. peneplana Berger, 1936 


Original combination. “Erebia aethiops ESPER, race 
peneplana, nov. race” Berger, 1936 Lambillionea 36: 199. 

Current combination. Erebia aethiops peneplana 
Berger, 1936. 

Current status. Valid subspecies. 

Type material. Paratypes 3¢6¢ (ZMH 835588-— 
835590) (Fig. 139). “PARATYPE / Collection / Luc. 
Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 5/8/1936 
/ Coll. Luc. BERGER.” // [blank label] // “ZMH835588”; 
“PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. 
Boyes) / BELGIQUE / 5/8/1936 / Coll. Luc. BERGER.” 


Evolutionary Systematics 5 2021, 193-261 243 


Ly pe 
Coen.corlana Hb... 
nba. hance. Bales 


ab.m fre as iia Bi uh. 
Oko Budbatek ZBL. A924. 
Evisa, Corse 


Juli 1921 
Col.0.Bubacek 


Evisa Corse 
Jind 1925 
| Col.O.Bubacek 


131a* 


[zm 327747| 


italia central. 
Mti.Simmbruini 


ae. | SEE 


Pers.Elbursgeb. 
Kendevanpass 
2800m,3,-8.7.36 
ee 


‘Coll, Bytinski-Salz: 
:Eing, Nr." 20 - 60: 


| ‘fu fi manasa 


7MH 827722| 


1cm 


Plate 19. 130. Coenonympha corinna ab. macrophthalmica Bubacek, 1923; 131. Coenonympha corinna ab. gynandra Bubacek, 
1923; 132. Coenonympha glycerion f. pulcherrima Warnecke, 1942; 133. Coenonympha glycerion ab. oikeia Dannehl, 1927; 
134. Coenonympha leander iranica Schwingenschuss, 1939; 135. Coenonympha oedippus rhenana Gradl, 1933. a. Dorsal view, 
b. ventral view. 


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244 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


// {blank label] // “ZMH835589”; “PARATYPE / Collec- 
tion / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE 
/ 16/8/1934 / Coll. Luc. BERGER.” // [blank label] // 
“ZMH835590”. 

Type locality. Belgium: “env. de Han-sur-Lesse”’. 


140. clorinda Hartig, 1940 


Original combination. “Erebia epiphron Fn. ssp. clorin- 
da n.” Hartig, 1940 Mem. Soc. Ent. Ital. 18: 186. 

Current combination. Erebia epiphron clorinda 
Hartig, 1940. 

Current status. Junior subjective synonym of Erebia 
epiphron amplevittata Verity, 1921. 

Type material. Paratypes 1619 (ZMH 827718- 
827719) (Fig. 140). “Paratypus 9 / Er. epiphr. / clorinda 
/ Hartig” // “1700 m / Marche / Bolognola / 20.VI 1937 
/ Querci” // [blank label] // “ZMH 827718”; “Paratypus 
9 / Er. epiphr. / clorinda / Hartig” // “1700 m / Marche / 
Bolognola / 20. VI 1937 / Querci” // [blank label | // “ZMH 
827718”; “Paratypus ¢ / Er. epiphr. / clorinda Hartig” // 
“1700 m / Marche / Bolognola / 20. VI 1937 / Querc1” // 
“Tblank label] // “ZMH 827719”. 

Type locality. Italy: “Bolognola”. 


141. vogesiaca Goltz, 1914 


Original combination. “Erebia epiphron vogesiaca” 
Goltz, 1914 Dt. Ent. Z. Iris 28:107—109. 

Current combination. Erebia epiphron vogesiaca 
Goltz, 1914. 

Current status. Junior subjective synonym of Erebia 
epiphron mackeri Fuchs, 1914). 

Type material. Syntype 14 (ZMH 827679) (Fig. 141). 
“COTYPE / vogesiaca / v.d.Goltz” // “VII 1913 / Faux- 
kopf / Vogesen” // “Fauxkopf / Juli 1913 / Kesenheimer” 
/1 “ZMH 827679”. 

Type locality. France: “Vogesen”’. 


142. leonhardi Fruhstorfer, 1918 


Original combination. “Evebia melas leonhardi sub- 
spec. nova” Fruhstorfer, 1918 Arch. Naturgesch. 84 
AP NS 1. 

Current combination. Erebia melas leonhardi 
Fruhstorfer, 1918. 

Current status. Valid subspecies. 

Type material. Syntypes 11¢9 (ZMH 827727- 
827737) (Fig. 142). “Velebit, Ostaria” // “Cotype” // 
“Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” 
/1 “ZMH 827727”; “Velebit, Ostaria” // “Cotype” // “Co- 
typus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // 
“ZMH 827728”, “Velebit, Ostaria” // “Cotype” // “Coty- 
pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827729”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 


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827730”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827731”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827732”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827733”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827734”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827735”; “Velebit, Ostaria” // “Cotype” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827736”; “Velebit, Ostaria” // “Cotypus” // “melas leon- 
hardi Cotype / Velebit Ostria 19[..]” // “Coll. Bytins- 
ki-Salz / Eing. Nr. 20, 1960” // “ZMH 827737”. 
Type locality. Croatia: “Velebit, Ostaria”’. 


143. paradisi Dannehl, 1929 


Original combination. “Erebia stygne O. paradisi Dhl.” 
Dannehl, 1929 Mitt. Munch. Ent. Ges. 19: 98. 

Current combination. Erebia meolans paradisi 
Dannehl, 1929. 

Current status. Junior subjective synonym of nomi- 
notypical Erebia meolans (de Prunner, 1798). 

Type material. Syntypes 3¢¢ (ZMH 835585-— 
835587) (Fig. 143). “19.6” / “Apenn. merid. / Mt. Par- 
adiso / 1500-2000 m / coll. F. Dannehl” // “19.6. 1928 / 
Mte. Paradiso / Central — Apenninen” // “ZMH 835585”; 
“22.6” / “Apenn. merid. / Mt. Paradiso / 1500-2000 m 
/ coll. F. Dannehl” // “20.6. 1928 / Mte. Paradiso / Cen- 
tral — Apenninen” // “ZMH 835586”; “19.6” / “Apenn. 
merid. / Mt. Paradiso / 1500-2000 m / coll. F. Dannehl” 
// “19.6. 1928 / Mte. Paradiso / Central — Apenninen” // 
“ZMH 835587”. 

Type locality. Italy: “Monte Paradiso 1600-1900 m, 
Monte Rotella, Montagna Grande 1800-2100 m (stil. 
Abruzzen).” 


144. benacensis Warren, 1933 


Original combination. “Erebia ottomana ssp. benacen- 
sis nov.” Warren, 1933 Ent. Rec. 45: 40. 

Current combination. Erebia ottomana benacensis 
Warren, 1933. 

Current status. Junior subjective synonym of nomi- 
notypical Erebia ottomana Herrich-Schaffer, (1847). 

Type material. Syntypes 5¢¢ (ZMH 833449- 
833453) (Fig. 144). “4.7” // “Teriolis merid. / Monte 
Baldo / 2000 m. / coll. F. Dannehl” // “4.7. 1930 / Mte. 
Baldo 2000 m / (Gardasee)” // “ZMH 833449”; “4.7” // 
“Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dan- 
nehl” // “4.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // 
“ZMH 833450”; “5.7” // “Teriolis merid. / Monte Baldo 
/ 2000 m. / coll. F. Dannehl” // “5.7. 1930 / Mte. Baldo 
2000 m / (Gardasee)” // “ZMH 833451”. 

Type locality. Italy: “Mte. Baldo near Lake Garda”. 


Evolutionary Systematics 5 2021, 193-261 245 


Litorale auatr. 
Triest Umgbg. 


« STAUDER 


a 
ins, 
— 
Lio} 


Eing. Nr. 5, 1949: 


[zm 827739 


ae 
Ubud. U... Motles 
| Flavofasciata 


Type 0 


Tellin. (R.d.Boyes.)_ 
BELGIQUE. 
£7. Ff /ABE. 

Coll-Lue. BERGER. 


ZMH 833469 


- Fauxkopf 
Julh1913 
Kesenheimer 


ZMH 327679] 


ZMH 827719| 


140b 


141b 
1cm 


Plate 20. 136. Coenonympha pamphilus galvagnii Stauder, 1923; 137. Coenonympha tullia f. maculata Warnecke, 1942; 138. Erebia 
flavofasciata Heyne, 1895; 139. Erebia aethiops peneplana Berger, 1936; 140. Erebia epiphron clorinda Hartig, 1940; 141. Erebia 
epiphron vogesiaca Goltz, 1914. a. Dorsal view, b. ventral view. 


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246 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


145. kotzschae Goltz, 1937 


Original combination. “Erebia kotzschae” Goltz, 1937 
Ent. Rdsch. 54: 363. 

Current combination. Paralasa kotzschae (Goltz, 
1937). 

Current status. Valid species. 

Type material. Syntype 146 (ZMH 827715) (Fig. 
145). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / 
Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. 
Kotzsch” // “COTYPE / kotzschae / v.d. Goltz” // “Kots- 
chi v.d. Goltz” // “Coll. Bytinski-Salz / Eing Nr 20 — 60” 
/1 “ZMH 827715”. 

Type locality. Afghanistan: Hindukush. 


146. maxima Bang-Haas, 1933 


Original combination. “Satyrus autonoe maxima O. B.- 
Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 98. 

Current combination. Hipparchia autonoe maxima 
(Bang-Haas, 1933). 

Current status. Valid subspecies. 

Type material. Syntypes 2¢¢ (ZMH 827681- 
827682) (Fig. 146). “Kansu mer. / Min-chan / Tanho / 
2500 m. Juli” // “Co-Type / O. Bang-Haas” // “L.13 / Sat. 
autonoé - / maxima / O.B.H.” // “Sammlung / R. Jahnig 
/ Eing Nr. 29, 1949” // “ZMH 827681”; “Kansu mer. or. 
/ Fukiang / Peilingschan sept. / 2500 m. Juli” // “Co- 
Type / O. Bang-Haas” // “L.14 / Sat. autonoé - / maxima 
/ O.B.H.” // “Sammlung / R. Jahnig / Eing Nr. 29, 1949” 
// “ZMH 827682”. 

Type locality. China: “Kansu mer., Lihsien, Kialing 
FluB, 2500 m.” 


147. persiscana Verity, 1937 


Original combination. “Eumenis allionii, G.-H. = fat- 
ua Freyer, persiscana, nom. nov.” Verity, 1937 Ent. Rec. 
49: 100. 

Current combination. Hipparchia fatua persiscana 
(Verity, 1937). 

Current status. Valid subspecies. 

Type material. Syntypes 31¢¢ (ZMH 827683- 
827713) (Fig. 147). “Iran / Keredj 1600 m / VIII 1936 / 
leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. 
Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827683”; 
“Tran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / 
coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz 
/ Eing. Nr. 20 - 60” // “ZMH 827684”; “Iran / Keredj 
1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytinski-Salz” 
// “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // 
“ZMH 827685”; “Iran / Keredj 1600 m/ VIII 1936 / leg. 
F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827686”; “Iran 
/ Keredj 1600 m/ VIII 1936 / leg. F. Brandt / coll. By- 
tinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. 


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Nr. 20 - 60” // “ZMH 827687”; “Iran / Keredj 1600 m / 
VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Coty- 
pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827688”; “Iran /Keredj 1600 m/ VIII 1936/leg. F. Brandt 
/ coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz 
/ Eimg. Nr. 20 - 60” // “ZMH 827689”; “Iran / Keredj 
1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytinski-Salz” 
// “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // 
“ZMH 827690”; “Iran / Keredj 1600 m / VIII 1936 / leg. 
F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827691”; “Tran 
/ Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. By- 
tinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. 
Nr. 20 - 60” // “ZMH 827692”, “Iran / Keredj 1800 m 
/ 21.8.1936 / Brandt” // “Cotypus” // “ssp. persicana / 
Vrty” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827693”; “Iran / Keredj 1600 m / VHI 1936 / leg. F. 
Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827694”; “Tran / 
Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytins- 
ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 
20 - 60” // “ZMH 827695”; “Iran / Keredj 1800 m / 25/8 
1936 / Brandt” //“Cotypus” // “Coll. Bytinski-Salz / Eing. 
Nr. 20 - 60” // “ZMH 827696”; “Iran / Keredj 1800 m / 
25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz 
/ Eing. Nr. 20 - 60” // “ZMH 827697”; “Iran / Keredj 
1600 m/ VU 1936 / leg. F. Brandt / coll. Bytinski-Salz” 
// “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // 
“ZMH 827698”; “Iran / Keredj 1600 m / VIII 1936 / leg. 
F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827699”; “Tran 
/ Keredj 1800 m / 25/8 1936 / Brandt” // “201” // “Coty- 
pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 
827700”; “Iran / Keredj 1800 m / 21/8 1936 / Brandt” // 
“Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // 
“ZMH 827701”; “Iran / Keredj 1200 m / 1936 / Brandt” 
// “1600 25/VIIT’ // “Cotypus” // “Coll. Bytinski-Salz / 
Eing. Nr. 20, 1960” // “ZMH 827702”; “Iran / Kered] 
1600 m/ VII 1936 / leg. F. Brandt / coll. Bytinski-Salz” 
// “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // 
“ZMH 827703”; “Iran / Keredj 1600 m / VIII 1936 / leg. 
F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827704”; “Tran / 
Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytins- 
ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 
20 - 60” // “ZMH 827705”; “Iran / Keredj (1200 m.) / 
1936 / Brandt” //“Cotypus” // “Coll. Bytinski-Salz / Eing. 
Nr. 20, 1960” // “ZMH 827706”; “Iran / Keredj 1600 m / 
25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz 
/ Eing. Nr. 20 - 60” // “ZMH 827707”; “Iran / Keredj 
1600 m/ VU 1936 / leg. F. Brandt / coll. Bytinski-Salz” 
// “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” 
// “ZMH 827708”; “Iran / Keredj 1600 m / VIII 1936 / 
leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. 
Bytinski-Salz / Eing. Nr. 20 - 60” //“ZMH 827709”; “Iran 
/ Keredj 1600 m/ VHI 1936 /leg. F. Brandt / coll. Bytins- 
ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 
20 - 60” // “ZMH 827710”; “Iran / Keredj 1200 m/ 1936 


Evolutionary Systematics 5 2021, 193-261 247 


Apenn, merid. 
Mt. Paradiso 
1500-2000 m 


Cotypus 


+ 


|. a 
ZMH 835585. 


erid, 
aldo 


Teriolis 


Monte Nord-Ost-Hindukusch 


Nuksan-Pass-Nordseite 
Alpenwiesenzone 
3500-4000m Mitte Juli 
leg. H.&E. Kotzsch 


[ou Bytinski-Salz: 
Eing Nr 20 - 69: 


f Sammlung 
R.Jaltnig 
Eing.Nr.29,1949 


Kansu mer. or. 
Fukiang , 
Peilingschan sept. 
2500 m. Juli 


Plate 21. 142. Erebia melas leonhardi Fruhstorfer, 1918; 143. Erebia meolans paradisi Dannehl, 1929; 144. Erebia ottomana 
benacensis Warren, 1933; 145. Paralasa kotzschae (Goltz, 1937); 146. Hipparchia autonoe maxima (Bang-Haas, 1933). a. Dorsal 
view, b. ventral view. 


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248 


/ Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 
20 - 60” // “ZMH 827711”; “Iran / Keredj 1600 m / VIII 
1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” 
// “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 
827712”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. 
Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- 
tinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827713”. 

Type locality. Iran: Tehran, “Keredj [Karaj], in the El- 
burz Mountains, at m. 1500 to 1600”. 


148. ab. pallida Bubacek, 1923 


Original combination. “Satyrus neomiris God. ab. nova 
pallida Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 
72: (30). 

Current combination. Hipparchia neomiris ab. pal- 
lida (Bubacek, 1923). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 1¢ (ZMH 
827726) (Fig. 148). “Type / Sat. neomiris God / ab. pal- 
lida Bub. / Otto Bubacek Z.BG. 1922” // “Col de Verg. 
Cor- / se Juli 1921 / Col. O. Bubacek” // [blank label] // 
“ZMH 827726”. 

Original locality. France: Corse, Col de Verg. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of S. neomiris (Godart, 1824). As stated by 
article 45.6.2 (ICZN 1999) it is deemed to be an infra- 
subspecific name (the author used “aberration”, “ab.’) 
and is hence unavailable. 


149. ab. albinervata Warnecke, 1944 


Original combination. “[Satyrus semele L.] n. ab. albin- 
ervata” Warnecke, 1944 Dt. Ent. Z. Iris 57: 40. 

Current combination. Hipparchia semele ab. albin- 
ervata (Warnecke, 1944). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Holotype” 19 
(ZMH 827749) (Fig. 149). “Insel Sylt / Kampen / VII 
1942” // “n.f. albinervata / Warnecke / Type” // “Holo- 
type” // “Sammlung G. Warnecke / Eing. Nr. 5, 1949” // 
“ZMH 827749”. 

Original locality. Germany: “Sylt, Kampen”. 

Remarks. Warnecke (1944) proposed this name as a 
form of S. semele (Linnaeus, 1758). As stated by article 
45.6.2 (ICZN 1999) it is deemed to be an infrasubspe- 
cific name (the author used “aberration”, “ab.”) and is 
hence unavailable. 


150. siviae Fuchs, 1954 


Original combination. “Lymanopoda shefteli siviae” 
Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84. 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Current combination. Lymanopoda shefteli siviae 
Fuchs, 1954. 

Current status. Valid subspecies. 

Type material. Lectotype 14 (ZMH 827764) and 3 
SS paralectotypes (ZMH 827765—827767) (Fig. 150). 
“Lymnopoda / siviae / v. Fuchs / Cordillere von / Siv- 
ia, Ayacucho / 2500 m.” // “Type” // “Holotype / Lym- 
nopoda siviae / von Fuchs” // “Lymnopoda galactea / 
shefteli Dyar / G. Lamas. det. 1978” // “Lectotype 3 
/ Lymnopoda / siviae Fuchs / by G. Lamas, 1978” // 
“Sammlung / W. v. Fuchs / Eing Nr. 4, 1956” // “ZMH 
827764”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // 
“285 / galactea ab. / Rio Piene 2600 m. 7/20” // “ZMH 
82765”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // 
“ZMH 82766”; “Rio Piene / ca. 2600 m / 7/1920 / 
Type” // “ZMH 82767”. 

Type locality. Peru: “... oberhalb Sivia gefangen, 
[above Sivia] 2500-3000 m”. 


(<4 


151. cocuzzana Stauder, 1914 


Original combination. “Melanargia arge Sulz. (=am- 
phitrite Hb.) cocuzzana m., subspec. nova” Stauder, 1914 
Zeit. Wiss. Ins. Biol. 10: 375, f. 12-15. 

Current combination. Melanargia arge cocuzzana 
Stauder, 1914. 

Current status. Junior subjective synonym of nomi- 
notypical Melanargia arge (Sulzer, 1776). 

Type material. Paratypes 3¢6¢ (ZMH 827673- 
827675) (Fig. 151). “Calabria / Umg. Paola-Co-senza 
4.6.1913 / H. Stauder” // “arge cocuzzana Stau- /der 
Paratype” // “ZMH 827673”; “Calabria / Umg. Paola- 
Co-senza 4.6.1913 / H. Stauder” // “cocuzzana” // “arge 
cocuzza- / na Stauder / Paratype” // “ZMH 827674”; “Ital. 
mer. / pen.-Surrentina / 8.6.1913 / H. Stauder” // “Arge 
cocuzzana / Stauder” // “arge cocuzzana Stau- / der Para- 
type” // “ZMH 827675”. 

Type locality. Italy: vicinity of Paola, Monte Cocuzzo 
“ an den Hangen [slopes of | des Monte Martinello und 
Cocuzzo bei Paola”. 


152. ab. huenei Hirschke, 1910 


Original combination. “Pararge hiera var. ominata 
Krul. ab. Huenei” Hirschke, 1910 Verh. Zool. Bot. Ges. 
Wien 60: 411. 

Current combination. Lasiommata petropolitana 
ominata ab. huenei (Hirschke, 1910). 

Current status. Infrasubspecific and hence unavail- 
able name.. 

Original material. Labelled as “Type” 1¢ (ZMH 
827717) (Fig. 152). “Estonia / Lechts / FR. V. HUENE” 
// “Type / Verh. z.b.g./ LX. 1910” // “Pararge hiera / var. 
ominata / ab. huenei / Hirschke / Estland” // “Coll. By- 
tinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827717”. 

Original locality. Estonia: “aus Esthland”. 


Evolutionary Systematics 5 2021, 193-261 249 


IRAN | aaa 
0 tinski-Salz |_- 
Bree Sern ZMH #27693 


Brandt 


92h 
Col. Oo. an an 


mange 827726 


E: Col de Nerg.Cor- 


Ry maojoda 
dovca 
2 ts Hechism 
Gorvcttes oct 
Jeaca, byac who 
hSt0m. 


HOLOTYPE 
Lyman pods. fini 
yon Fuchs 


LECTOTYr! 
Lymane 
pas Pocus 


by 6 Lomas, 1478 


ong 


Vi Ta 


“Sammlung: 
6. Beare t 


L gmanapoels galacts. a 
craseels Ager 


G.Lamss. det. 1978 


ee tung 
W.'v.¢Euchs 
Eing. ‘Ne, 1956; 


ai PS 


Calabria 
| Umg.Paola-Co- 
— t& ‘a, 6. 19h 3 
| Stauder 9 


ren 


o 


zm 827674] 


Plate 22. 147. Hipparchia fatua persiscana (Verity, 1937); 148. Hipparchia neomiris ab. pallida (Bubacek, 1923); 149. Hipparchia 
semele ab. albinervata (Warnecke, 1944); 150. Lymanopoda shefteli siviae Fuchs, 1954; 151. Melanargia arge cocuzzana Stauder, 


1914. a. Dorsal view, b. ventral view. 


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250 


Remarks. Hirschke (1910) proposed this name as an 
aberration of P. h. ominata Krulikowsky, 1903, therefore, 
as stated by articles 45.6.1 and 45.6.2 ICZN 1999) it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 


153. felix Warnecke, 1929 


Original combination. “Pararge felix n. spec. Warn.” 
Warnecke, 1929 Int. Ent. Z. 22: 365. 

Current combination. Lasiommata felix (War- 
necke, 1929). 

Current status. Valid species. 

Type material. Paratype 12 (ZMH _ 827721) 
(Fig. 153). “S.W. Arabia / Yemen, Sanaa / II.1928. Ex- 
pedi- / tion Rathjens” // “28.2. /Sanaa” // “Pararge felix / 
Warnecke 1929 / 9 Paratype” // “Para-Type” // “Pararge 
maera - | felix Warn.” // “Sig. G. Warnecke / Eing. Nr. 5, 
1949” // “ZMH 827721”. 

Type locality. “Yemen, Sanaa”. 


154. ab. diluta Bubacek, 1923 


Original combination. “Pararge aegeria L. ab. nova diluta 
Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30). 

Current combination. Pararge aegeria ab. diluta 
Bubacek, 1923. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
827678) (Fig. 154) “Type / Par. aegeria L. / ab. diluta 
Bub. / Otto Bubacek ZBG 1922” // [blank label] // “Evisa, 
Corse / Juli 1921 / Col. O. Bubacek” // “ZMH 827678”. 

Original locality. France: Korsika [Corse]. 

Remarks. Bubacek (1923) proposed this name as an 
aberration of P. aegeria (Linnaeus, 1758), therefore, as 
stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. 


155. f. albiplaga Warnecke, 1942 


Original combination. “Pararge algeria L. subsp. 
egerides Stdgr. n. f. albiplaga 9°.” Warnecke, 1942 Dt. 
Ent. Z. Iris 56: 102. 

Current combination. Pararge aegeria egerides f. 
albiplaga Warnecke, 1942. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 19 (ZMH 
827668) (Fig. 155). “Holstein / Segeberg / 2.-10.8.35” // 
“nf. albiplaga / Warn. Type / Iris 1942, 102” // “Type” // 
“ZMH 827668”. 

Original locality. Germany: Holstein [label indicates 
Segeberg]. 


evolsyst.pensoft.net 


Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Remarks. Warnecke (1942) proposed this name as a 
form of P. a. egerides (Staudinger, 1871), therefore, as 
stated by article 45.6.1 it is deemed to be an infrasubspe- 
cific name (the author expressly gave it infrasubspecific 
rank) and is hence unavailable. In the original descrip- 
tion the name “aegeria” is wrongly spelled “algeria” (M. 
Wiemers pers. comm. ). 


156. ab. hermini Hirschke, 1910 


Original combination. “Pararge megera var. lyssa B. 
ab. Hermini” Hirschke, 1910 Verh. Zool. Bot. Ges. Wien 
60: 411. 

Current combination. Lasiommata megera lyssa ab. 
hermini (Hirschke, 1910). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 1¢ (ZMH 
827720) (Fig. 156). “Hirschke.Wien / Zara / Mai” // “Type 
/ Verh. Z.G.B. / LX, 1910” // “Pararge megera / var. Ly- 
ssa ab. / hermini Hirschke / Dalmatien 5.1909” // “Coll. 
Bytinski-Salz / Eing Nr 20 — 60” // “ZMH 827720”. 

Original locality. Croatia: Dalmatia, vicinity of Gra- 
vosa and Zara. 

Remarks. Hirschke (1910) proposed this name as an 
aberration of P. m. lyssa “B.” (sic, recte Hubner, 1829), 
therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 
1999) it is deemed to be an infrasubspecific name (the 
author used “aberration”, “ab.”, or the author expressly 
gave it infrasubspecific rank) and is hence unavailable. 


157. picota Fuchs, 1954 


Original combination. “Argyrophorus picota” Fuchs, 
1954 in Titschak, Beit. zur Fauna Perus 4: 84. 

Current combination. Punargentus angusta picota 
(Fuchs, 1954). 

Current status. Valid subspecies. 

Type material. Lectotype 14 (ZMH 827750) (Fig. 157) 
7319 paralectotypes (ZMH 827751-827758). “Argyro- 
phorus / picota 3 / y. Fuchs / Cordillere von / Ayacucho 
/ 4500 m” // “Punargentus lamna / angusta Weymer / det. 
W. Heinrich 3” // “Type” // “Lectotype ¢ / Argyrophorus 
/ picota Fuchs / by G. Lamas, 1978” // “Sammlung / W. 
v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827750”; “Punar- 
gentus lamna / angusta Weymer / det. W. Heinrich 3” // 
Argyrophorus | picota 3 / v. Fuchs / Cordillere von / Ay- 
acucho / 4500 m” // “Type” // “Sammlung / W. v. Fuchs / 
Eing. Nr. 4, 1956” // “ZMH 827751”; “Ica / Ayacucho / ca. 
4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta 
Weymer / det. W. Heinrich 3” // Coll. H. Rédinger / Eing. 
Nr. 30, 52” // ZMH 827752”; “Ica / Ayacucho / ca. 4000 m. 
/ 7/1920 / Type” // “Punargentus lamna / angusta Weymer / 
det. W. Heinrich 3” // Coll. H. Rédinger / Eing. Nr. 30, 52” 
/1 ZMH 827753”, “Ica / Ayacucho / ca. 4000 m. / 7/1920 / 
Type” // “Punargentus lamna / angusta Weymer / det. W. 
Heinrich @” // “133 / angusta 8/20 / Ayacucho 4/5000 m” 


Evolutionary Systematics 5 2021, 193-261 251 


LW, brabs 
ies ina 

1914. Et pede~ 
Him Fates cus 


Porarce hiera 
Var. oa . n Ve 
‘hutnei 
Est lineal’ rsthiee 


Coll, Bytingki-Salz: 
‘Eing, Nr., 20 - 69: 


Beosgextislenme 
nc STE Weyer 


oez: 7. eirmthé 


rage mewe 
CFE, 

ver Lyssa ag 

Baring i'ma 
almatien F.71999 


157b 


11cm 


Plate 23. 152. Lasiommata petropolitana ominata ab. huenei (Hirschke, 1910); 153. Lasiommata felix (Warnecke, 1929); 
154. Pararge aegeria ab. diluta Bubacek, 1923; 155. Pararge aegeria egerides f. albiplaga Warnecke, 1942; 156. Lasiommata 
megera lyssa ab. hermini (Hirschke, 1910); 157. Punargentus angusta picota (Fuchs, 1954). a. Dorsal view, b. ventral view. 


evolsyst.pensoft.net 


252) Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


// “Coll. H. Rodinger / Eing. Nr. 30, 52” // ZMH 827754”: 
“Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- 
gentus lamna / angusta Weymer / det. W. Heinrich 3” // 
Coll. H. Rédinger / Eing. Nr. 30, 52” // ZMH 827756”; 
“Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- 
gentus lamna / angusta Weymer / det. W. Heinrich 3” // 
Coll. H. Rédinger / Eing. Nr. 30, 52” // ZMH 827757”; 
“Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- 
gentus lamna / angusta Weymer / det. W. Heinrich 9” // 
Coll. H. Rodinger / Eing. Nr. 30, 52” // ZMH 827758”. 

Type locality. Peru: near Ayacucho “... Kodillere bei 
Ayacucho, nur in Hohen tiber 4000 m, bis 5000 m” [Cor- 
dillere close to Ayacucho, only at altitudes above 4000 m, 
to 5000 m]. 


158. arminii Stauder, 1916 


Original combination. “Epinephele ida arminii subsp. 
nova.” Stauder, 1916 Zeit. Wiss. Ins. Biol. 12: 62. 

Current combination. Pyronia cecilia arminii 
(Stauder, 1916). 

Current status. Junior subjective synonym of Pyronia 
cecilia cecilia (Vallantin, 1894). 

Type material. Syntype 14 (ZMH _ 827716) 
(Fig. 158). “Type” // “Calabria mer. / Aspromonte 1600 
m. / 26.6.1920 / H. Stauder legit.” // “Ida Arminii / Staud- 
er” // “Type” // “Sammlung / G. Warnecke / Eing. Nr. 5, 
1949” // “ZMH Nr. 5, 1949”, 

Type locality. Italy: Calabria. 


159. liupiuschani Bang-Haas, 1933 


Original combination. “Satyrus actaea liupiuschani O. 
B.-Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 99. 

Current combination. Satyrus ferula liupiuschani 
Bang-Haas, 1933. 

Current status. Junior subjective synonym of Satyrus 
ferula ganssuensis Grum-Grshimailo, 1893. 

Type material. Syntype 14 (ZMH 827669) (Fig. 
159). “Kansu mer. / Tsing-schui / Liu-pin-schan / 2000 
m. Juli” // “Co-Type / O. Bang-Haas” // “L.17 / Sat. ac- 
taea - / liupiuschani O.B.H.” // “Sammlung / R. Jahnig / 
Eing. Nr.29,1949” // “ZMH 827669”. 

Type locality. China: “Kansu mer., Tsingschui, Liu 
Piu Schan, 2000 m”. 


160. kintschouensis Bang-Haas, 1939 


Original combination. “7riphysa phryne kintschouensis 
O. Bang-Haas, subsp. nov.” Bang-Haas, 1939 Dt. Ent. Z. 
Ins S3"53" 

Current combination. Coenonympha phryne kint- 
schouensis (Bang-Haas, 1939). 

Current status. Junior subjective synonym of Co- 
enonympha dohrnii nervosa Motschulsky, 1866. 


evolsyst.pensoft.net 


Type material. Syntypes 2¢¢ (ZMH 827759- 
827760) (Fig. 160). “18.” // “Mandschuria mer. occ. / 
Prov. Fengtien / Kintschou / 100 m. April” // “Co-Type / 
O. Bang-Haas” // “Phryne / v. kintschouensis / O.B.H.” 
// “L.21 / Triphysa phyrne - / kintschouensis / O.B.- 
Haas” // “Sammlung R. Jahnig / Eing Nr. 29, 1949” // 
“ZMH 827759”; “17.” // “Mandschuria mer. occ. / Prov. 
Fengtien / Kintschou // “Co-Type / O. Bang-Haas” // 
“L,.22 / Triphysa phyrne - / kintschouensis / O.B.-Haas” 
// “Sammlung R. Jahnig / Eing Nr. 29, 1949” // “ZMH 
827760”. 

Type locality. China: “Manchuria merid., Prov. Feng- 
tien, Kintschou, 100 m.”. 


Unpublished, unavailable names 


161. ab. minor Meinhard 


Original combination. Melanargia galathea v. sakaria 
ab. minor Meinhard. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Type” 1¢ (ZMH 
835597) (Fig. 161). “Type” // 7/7 1920 / Gottingen / 
Warteberg” // “ZMH 835597”. 

Remarks. This taxon cannot be found in the literature. 
As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is 
deemed to be an infrasubspecific name (the author used 
“aberration”, “ab.”, or the author expressly gave it infra- 
subspecific rank) and is hence unavailable. 


162. ab. caeca Dannehl 


Original combination. Epinephele ida ab. caeca Dan- 
nehl. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Cotype” 14 (ZMH 
833467) (Fig. 162). “1.6” // Sicilia / Mt. Cuccio / coll. 
Dannehl” // “Cotype / caeca / Dhl” // “Cotype.” // “1.6 
1932 / Mt. Cuccio, / -Sizilien-” // “ZMH833467”. 

Remarks. This taxon cannot be found in the literature. 
As stated by article 45.6.2 (ICZN 1999) it is deemed to be 
an infrasubspecific name (the author used “aberration”, 
“ab.”) and is hence unavailable. The current name of the 
species 1s Pyronia cecilia (Vallantin, 1894). 


163. achaltekkensis Holik 


Original combination. Satyrus actaea achaltekkensis 
Holik. 

Current status. “in litteris’ name and hence not 
available. 

Original material. Labelled as “Cotypes” 24'3' (ZMH 
827670—-827671) (Fig. 163). “Transcaspia / Jablonowka / 
Achal Tekke / 2000 m. Juli” // “Co-Type” // “L.18 / Sat. 


Evolutionary Systematics 5 2021, 193-261 253 


Kansu mer- 
Tsing-schui 
‘Liu-pin-schan! 
2000 m. Juli: 


158a 


' Sammlung 
: RJahhig | 
|Eing.Nr.29,1949 | 


158b 


Mandschuria mer.oce, 
rov. Fengtien 


Kintgthou 
(100m ll 


“Sammlung 
 RJabnig 5 
Fina Nr.29,1949 : 


> = <= 2MH_835597 


Transcaspia 
Jablonowka 
Achal Tekke 
2000m, Juli 


Plate 24. 158. Pyronia cecilia arminii (Stauder, 1916); 159. Satvrus ferula liupiuschani Bang-Haas, 1933; 160. Coenonympha 
phryne kintschouensis (Bang-Haas, 1939); 161. Melanargia galathea v. sakaria ab. minor Meinhard; 162. Epinephele ida ab. caeca 
Dannehl; 163. Satyrus actaea achaltekkensis Holik. a. Dorsal view, b. ventral view. 


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254 


actaea - | achaltekkensis Hol.” // “Sammlung / R. Jahnig 
/ Eing. Nr. 29, 1949” // “ZMH 827670”; “Transcaspia / 
Jablonowka / Achal Tekke / 2000 m. Juli” // “Co-Type” // 
“L.19/ Sat. actaea - / achaltekkensis Hol.” // “Sammlung 
/R. Jahnig / Eing. Nr. 29, 1949” // “ZMH 827671”. 

Original locality. Turkmenistan, Achal Tekke. 

Remarks. This taxon cannot be found in the litera- 
ture. The current name is Satyrus iranica daubi Gross & 
Ebett 1975: 


164. rossarmenia Bang-Haas 


Original combination. Satyrus actaea rossarmenia 
Bang-Haas. 

Current status. “in litteris” name and hence not 
available. 

Original material. Labelled as “Cotype” 14 (ZMH 
827672) (Fig. 164). “Kagysman / Russ. Armen” // “Co- 
Type” //“L.16 / Sat. actaea / rossarmenia ?” // “Sammlu- 
ng R. Jahnig / Eing. Nr.29, 1949” // “ZMH 827672”. 

Remarks. This taxon cannot be found in the literature. 
The current name is Satyrus amasina Staudinger, 1861. 


Family Riodinidae Grote, 1895 
Subfamily Nemeobiinae Bates 1868 
Unpublished, unavailable names 


165. ab. friesi Meinhard 


Original combination. Hamearis lucina ab. friesi 
Meinhard. 

Current status. “in litteris” name and hence not 
available. 

Original material. Labelled as “Type” 14 (ZMH 
835594) (Fig. 165). “Type” // “ab. friesi Mein.” // “10/5 
1911 / Gottingen / Hainberg” // “ZMH 835594”. 

Remarks. This taxon cannot be found in the literature. 
As stated by article 45.6.2 (ICZN 1999) it is deemed to 
be an infrasubspecific name (the author used “aberra- 
tion’, “ab.”) and is hence unavailable. The current name 
is Hamearis lucina (Linnaeus, 1758). 


Family Hesperiidae Latreille, 1809 
Subfamily Pyrginae Burmeister, 1878 
Tribe Carcharodini Verity, 1940 


166. wissmanni Warnecke, 1934 


Original combination. “Carcharodus swinhoei Wats. 
nov. subsp. Wissmanni Warn.” Warnecke, 1934 Mitt. 
Munch. Ent. Ges. 24: 20. 

Current combination. 
manni Warnecke, 1934. 

Current status. Valid subspecies. 

Type material. Paratypes 5<¢'¢' (ZMH 827637-ZMH 
827638, ZMH 827640—-ZMH 827642) (Fig. 166). “S. W. 
Arabien / Sana’a (Yemen) / 27.2.28 v. Wissm.” // “Sig. G. 


Carcharodus alceae wiss- 


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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Warnecke / Eing. Nr. 5, 1949” // “Carcharodus | Swin- 
hoei Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. 
Warnecke / Eing. Nr. 5, 1949” //“ZMH 827637”; “Alceae 
Swinhoei Wats. / S.W. Arabia / Sana’a (Yemen) 27.2.28 
/ Rathjens - v. Wissmann” // “Carcharodus | Swinhoei 
Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. War- 
necke / Eing. Nr. 5, 1949” // “ZMH 827638”, “Stid-Ara- 
bien / Yemen: San’aa / 8.8.31 Rathjens” // “Paratype” // 
Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827640”; 
“Stdwestarabien / Yemen: San’aa / Rathjens 24.6.31” 
// “Paratype” // Coll. Warnecke / Eing. Nr. 5, 1949” // 
“ZMH 827641”; “Stidwestarabien / Yemen: San’aa / 
Rathjens 24.7.31” // “Paratype” // “ZMH 827642”. 

Type locality. Yemen: Sanaa “Stdwest-Arabien, Sa- 
na’a (Yemen)” 


167. v. postesseloides Verity, 1938 


Original combination. “Spialia orbifer, Hub., race tes- 
selloides, H.-S., II gen. postesseloides, nom. nov.” Verity, 
1938 Ent. Rec. 50 Suppl. (4). 

Current combination. Spialia orbifer orbifer v. post- 
esseloides Verity, 1938. 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 443 (ZMH 
833498—-833501) (Fig. 167). “postesseloides Vrty” // 
“Macedonia / Olympus, 2500’ / Aug 24, 1935 / Romei” // 
[blank label] // “ZMH 833498”; “postesseloides Vrty” // 
“Macedonia / Olympus, 2500’ / July 25, 1936 / Rome” // 
[blank label] // “ZMH 833499”; “postesseloides Vrty” // 
“Macedonia / Olympus, 2500’ / July 6, 1936 / Romei” // 
[blank label] // “ZMH 833500”; “postesseloides Vrty” // 
“Macedonia / Olympus, 2000’ / July 8, 1936 / Romei” // 
[blank label] // “ZMH 833501”. 

Original locality. Greece: vicinity of Skala. 

Remarks. Verity (1938) proposed this name as “nom. 
nov.“ of S. orbifer tesselloides (Herrich-Schaffer, 1845), 
therefore, as stated by article 45.6.1 it is deemed to be 
an infrasubspecific name (the author expressly gave it 
infrasubspecific rank) and is hence unavailable. Spialia 
orbifer tesselloides is currently regarded as a junior sub- 
jective synonym of the nominate subspecies. 


Tribe Pyrgini Burmeister, 1878 


168. centralitaliae Verity, 1920 


Original combination. “Hesperia alveus, Hub., race 
centralitaliae, mihi” Verity, 1920 Ent. Rec. 32: 4. 

Current combination. Pyrgus alveus centralitaliae 
(Verity, 1920). 

Current status. Valid subspecies. 

Original material. Labelled as “Type” 433 
(ZMH 833502—833505) (Fig. 168). “centralitaliae 
Vrty” // “Marche / Bolognola / 6.VII.1937 / Quer- 
ci” // [blank label] // “ZMH 833502”; “centralitaliae 


Evolutionary Systematics 5 2021, 193-261 


Vrty” // “Marche / Bolognola / 13.VII.1937 / Quer- 
ci” // [blank label] // “ZMH 833503”; “centralitaliae 
Vrty” // “Marche / Bolognola / 22. VI.1937 / Querci” // 
[blank label] // “ZMH 833504”; “centralitaliae Vrty” // 
“Marche / Bolognola / 27.VI.1937 / Querci” // [blank 
label] // “ZMH 833505”. 

Original locality. Italy: Bologna. 

Remarks. Verity (1920) proposed this name as a 
“race” of H. alveus Hiibner, [1803]. As it was given in 
trinomy and the content of the description clearly indi- 
cates that a geographical race was meant, this name 1s 
available as subspecific (article 45.6 ICZN 1999). The 
year on the specimen labels (1937) indicate specimens 
were collected 17 years after description (1920) from 
the same location the original holotype material was 
collected. As a result, the black label of the specimens 
treated as “types” is erroneous. The specimen collect- 
ed from the same geographical location from which the 
type specimen was collected is called topotype. Accord- 
ing to ICZN (1999), a topotype has no formal standing 
and is not regulated by the Code. Therefore, these four 
specimens are erroneously labelled as types, should be 
annotated as “Not a Type” and are treated here as “Non- 
type” specimens 


169. badachschana Alberti, 1939 


Original combination. “Hesperia badachschana nov. 
sp.” Alberti, 1939 Ent. Rdsch. 56: 107. 

Current combination. Pyrgus badachschana (AI- 
berti, 1939). 

Current status. Valid species. 

Type material. Holotype 1 ¢ (ZMH 827639) (Fig. 169). 
“HOLOTYPE < / Hesperia / badachschana” |/ “Bad- 
achschan / Sebal..|-Tal / Alpenwiesenzone / 2800-3000 m 
Mitte Juni / leg. H.&E. Kotzsch” // “Coll. Bytinski-Salz 
Coll / Eing. Nr. 20, 1960 Eing” // “ZMH 827639”. 

Type locality. Afghanistan: Badakhshan. 

Remarks. We follow SEL declaration on gender 
agreement (Sommerer 2002), as a result it should stay 
as badachschana. 


170. picena Verity, 1920 


Original combination. “Hesperia foulquieri, Obth., race 
picena, mihi” Verity, 1920 Ent. Rec. 32: 4. 

Current combination. Pyrgus foulquieri picena 
(Verity, 1920). 

Current status. Valid subspecies. 

Original material. Labelled as “Type” 44'3' (ZMH 
833506—833509) (Fig. 170). “Picena Vrty” // “Marche / 
Bolognola / 2 VII 1937 / Querci” // [blank label] // “ZMH 
833506”; “Picena Vrty” // “Marche / Bolognola / 8 Ag 
1937 / Querci” // [blank label] // “ZMH 833507”; “Pi- 
cena Vrty” // “Marche / Bolognola / 8 Ag 1937 / Quer- 
ci” // [blank label] // “ZMH 833508”; “Picena Vrty” // 


255 


“Marche / Bolognola / 12 Ag 1937 / Querci” // [blank 
label] // “ZMH 833509”. 

Original locality. Italy: Bolognola, Sibillini Mts. 

Remarks. Verity (1920) proposed this name as a 
“race” of H. foulquieri Oberthur, 1910. As it was given 
in trinomy and the content of the description clearly in- 
dicates that a geographical race was meant, this name 1s 
available as subspecific (article 45.6 ICZN 1999). The 
year on the specimen labels (1937) indicate specimens 
were collected 17 years after description (1920) from 
the same location the original holotype material was 
collected. As a result, the black label of the specimens 
treated as “types” is erroneous. The specimen collect- 
ed from the same geographical location from which the 
type specimen was collected is called topotype. Accord- 
ing to ICZN (1999), a topotype has no formal standing 
and is not regulated by the Code. Therefore, these four 
specimens are erroneously labelled as types, should be 
annotated as “Not a Type” and are treated here as “Non- 
type” specimens. 


171. ab. alboradiata Bubacek, 1926 


Original combination. “Hesperia sidae Esp. var. on- 
opordi H.-S. ab. nova alboradiata Bub.” Bubacek, 1926 
Verh. Zool. Bot. Ges. Wien 76: 174. 

Current combination. Pyrgus sidae ab. alboradiata 
(Bubacek, 1926). 

Current status. Infrasubspecific and hence unavail- 
able name. 

Original material. Labelled as “Type” 14 (ZMH 
833497) (Fig. 171). “Type / H. sidae v. onopordi / ab. 
alboradiata Bub. / Z. B. G. 1926 O. Bubacek” // “Ttalia / 
Alassio — Liguria / 7/6 1925 / Coll. O. Bubacek” // [blank 
label] // “ZMH 833497”. 

Original locality. Italy: “Alassio, Liguria”. 

Remarks. Bubacek (1926) proposed this name as 
an aberration of H. sidae v. onopordi “H.-S.” (sic, recte 
Rambur, 1839), therefore, as stated by articles 45.6.1 and 
45.6.2 (ICZN 1999) it is deemed to be an infrasubspecif- 
ic name (the author used “aberration”, “ab.”, or the au- 
thor expressly gave it infrasubspecific rank) and is hence 
unavailable. P onopordi Rambur is a distinct species 
from P. sidae Esper, and both species occur sympatrical- 
ly in Liguria. 


Discussion 


With more than 2,000 type specimens of butterflies and 
moths (Lepidoptera), the Zoological Museum of Ham- 
burg (ZMH) is one of the most important natural histo- 
ry archives in Germany. In this catalogue, a total of 414 
specimens labelled as “types” belonging to the superfam- 
ily Papilionoidea are listed and discussed. This includes 
23 specimens of 16 “in litteris” (= unavailable) names, 43 
topotypes (non-type specimens), 97 original specimens 


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256 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


/ Pe bP ecccta/ 
22. Baty. Vira. 


| “Warnacka 


faran 


5 ce Lene 
ZMH 833500 


es 


> Coll. Warnecke | 
ZEing. Nr. 5, 1949 


Coll Bytinski-SalzColl 
Eing. Nr. 20, 1960Eing. 


Plate 25. 164. Satyrus actaea rossarmenia Bang-Haas; 165. Hamearis lucina ab. friesi Meinhard; 166. Carcharodus alceae 
wissmanni Warnecke, 1934; 167. Spialia orbifer orbifer v. postesseloides Verity, 1938; 168. Pyrgus alveus centralitaliae (Verity, 
1920); 169. Pyrgus badachschana (Alberti, 1939); 170. Pyrgus foulquieri picena (Verity, 1920); 171. Pyrgus sidae ab. alboradiata 
(Bubacek, 1926). a. Dorsal view, b. ventral view. 


of invalid infrasubspecific entities under the ICZN code, subspecies (representing 74 subspecies: 44 valid names 
and 251 type specimens (Fig. 172; Suppl. material 2: Ta- and 30 synonyms) (Suppl. material 1: Table S1, Suppl. 
ble S2). Out of 251 type specimens, 18 specimens repre- material 2: S2: Fig. 172). Of these 414 specimens, 171 are 
sent species (nine valid species), and 233 specimens are primary types (8 holotypes/lectotypes and 163 syntypes), 


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Evolutionary Systematics 5 2021, 193-261 257 


200 


oe 


100 
: 
Lo 
* 
, Paratype % 
50 be | 
pera 
Holotype oy 23 
ia 
Primary types Secondary types Original speci 
ae ill 


Figure 172. Type status among 414 “type” specimens of the lepidopteran superfamily Papilionoidea deposited in the Zoological 
Museum of Hamburg (ZMH). 


GEOGRAPHICAL DISTRIBUTION: OF TYPE LOCALITIES 


Bulgaria ma 5 
a Bo 


lran/Austria aol 8 


Afghanistan 


France 
China 
Germany 


Italy 


Figure 173. Geographical distribution of “type” localities among 171 representative “type” specimens of the lepidopteran super- 
family Papilionoidea deposited in the Zoological Museum of Hamburg (ZMH). Category A (six type localities) includes Algeria, 
Greece, Peru and Russia; category B (three type localities) includes Switzerland, Turkey and Spain; category C (two type localities) 
includes Armenia, Croatia and Yemen; category D (one type locality) includes Turkmenistan, Sweden, Romania, Macedonia. Leb- 
anon, Japan, Estonia, Czech Republic, Bosnia and Herzegovina, and Belgium. 


80 are secondary types, 120 are considered of invalid The comparison of our updated catalogue to the type 
infrasubspecific rank (original specimens), and 43 speci- catalogue of Weidner (1974) reveals 95 new additions 
mens are re-labelled as “Non-type” specimens (topotype) (237 specimens) to the type collection and type mate- 
(Fig. 172). rial of one herspertid species (1.e., Adopaea pfeifferi 


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258 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea 


Bytinski-Salz & Brandt, 1937) being potentially lost 
(one specimen) (Suppl. material 1: Tables S1, Suppl. 
material 2: Table S2). The latter species is currently a 
junior subjective synonym of Thymelicus hyrax (Leder- 
er, 1861). New additions to the type collection since 
the Weidner catalogue (1974) do not necessarily reflect 
“new type additions” in all instances. Except for a few 
“type” specimens donated after 1974 to the ZMH col- 
lection (e.g., from the former collection of the Altona 
museum), many specimens were already during Weid- 
ner’s time in the collection, but were either misidenti- 
fied or misplaced. Details of the changes and updates 
including the new types, changes to the type status, and 
changes to the specimen counts since the last catalogue 
of Weidner (1974) were summarized in Suppl. material 
1: Tables S1, Suppl. material 2: S2. 

Five countries (Italy — 37, Germany — 25, China — 16, 
France — 13, and Afghanistan — 10) represent the majority 
of type localities for 60% (101 of 172 “names”) of the 
Papilionoidea type collection followed by Iran and Austria 
(each represented by 8 “type” localities), Russia, Greece, 
Peru and Algeria (each with 6 “type” localities) (Fig. 173). 
Most type specimens come from the collection of Dr. 
George Warnecke, mainly from Europe and the Middle East 
(52 taxa), followed by the collection of Dr. Hanan Bytinsk1- 
Salz (20 taxa). The oldest type specimen of Papilionoidea 
at ZMH is the lectotype of Colias aurorina ab. crocopepla 
Warnecke, 1912 collected in Amur in 1887. The oldest 
described types are Parnassius mnemosyne giganteus 
Staudinger, 1886 and Parnassius nomion nominulus 
Staudinger, 1894, both are valid subspecies names from the 
collections of H. Rédinger and G. Warnecke, respectively, 
followed by Erebia flavofasciata Heyne, 1895, a valid 
species from the collection of von Nolte. 


Acknowledgements 


This project was funded by the CeNak “Digitalisierung 
der CeNak-Sammlungen’’. We also like to thank HASPA 
Lotterie Sparen for funding boxes and the City of Ham- 
burg for further funding via Bussgeldfonds. We thank the 
Gesellschaft der Freunde und Forderer des Zoologischen 
Museums Hamburg for providing financial support to the 
collections. The authors also acknowledge the members 
of the other Zoological Collections at the CeNak for their 
patience during our intensive photography sessions. Com- 
ments by Vladimir Lukhtanov (St. Petersburg, Russia), 
Niklas Wahlberg (Lund, Sweden), Alberto Zilli (London, 
UK), Karen McLachlan Hamilton (Ottawa, Canada) and 
Danilo Harms (Hamburg, Germany) substantially im- 
proved the manuscript. 


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Supplementary material | 


Table SI 


Authors: Reza Zahiri, Vazrick Nazari, Hossein Rajaei, 
Martin Wiemers, Maryam Fatahi, Matthias Seidel, 
Thure Dalsgaard, Martin Husemann 

Data type: excel table 

Explanation note: List of the 171 representative “type” 
specimens of the lepidopteran superfamily Papilion- 
oidea deposited in the Zoological Museum of Ham- 
burg (ZMH). 

Copyright notice: This dataset is made available under 
the Open Database License (http://opendatacommons. 
org/licenses/odbl/1.0). The Open Database License 
(ODbL) is a license agreement intended to allow us- 
ers to freely share, modify, and use this Dataset while 
maintaining this same freedom for others, provided 
that the original source and author(s) are credited. 

Link: https://do1.org/10.3897/evolsyst.5.63435 suppl 1 


Supplementary material 2 


Table S2 


Authors: Reza Zahiri, Vazrick Nazari, Hossein Rajaei, 
Martin Wiemers, Maryam Fatahi, Matthias Seidel, 
Thure Dalsgaard, Martin Husemann 

Data type: excel table 

Explanation note: List of the 414 “type” specimens of the 
lepidopteran superfamily Papilionoidea deposited in 
the Zoological Museum of Hamburg (ZMH). 

Copyright notice: This dataset is made available under 
the Open Database License (http://opendatacom- 
mons.org/licenses/odbl/1.0). The Open Database 
License (ODbL) is a license agreement intended to 
allow users to freely share, modify, and use this Data- 
set while maintaining this same freedom for others, 
provided that the original source and author(s) are 
credited. 

Link: https://do1.org/10.3897/evolsyst.5.63435 .suppl2 


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